Forest Wildlife Richard H. Yahner, Carolyn G. Mahan, and Amanda D. Rodewald

What is a Forest? ► Group of trees in defined area ► Geography – dependent  Coniferous forests (Western) ► Examples: Pacific coast Douglas fir, Rocky mountain lodgepole pine  Deciduous forests (Eastern) ► Examples: Lake States Aspen/birch, Central Mountains oak/hickory

History of Forests ► Pre-European forest covered 2/3 of North America  Disturbance regime: wildfires, native Americans ► 17 th century – 21 st century  53% decline in forest coverage  Timber harvesting, non-native pathogens (e.g., Chestnut blight, agricultural clearing ► Loss of all but 3-5% old growth forest ► Second growth forest dominates today

Forest Management and Ownership ► Commercial, reserved, and noncommercial forestlands—focuses on timber resources ► Forests used by 80-90% of N. American vertebrates ► Ownership of forests effects management decisions for wildlife  East—dominated by private  West – dominated by public (e.g., US Forest Service)

Early Legislation ► Forest Reserve Act (1891)  Created first forest reserves ► Organic Act (1897)  Established National Forest Service (later, USFS)  Theodore Roosevelt and Gifford Pinchot (1 st Head, USFS) set aside millions of acres ► Weeks Law (1911)  Created state and national partnerships for forest land acquisition

Other Legislation ► Multiple Use and Sustained Yield Act (1960)  Forests may be managed for wildlife and other uses ► National Environmental Protection Act (1969)  Environmental Impact Statements required ► Forest Rangeland Renewable Resources Planning Act (1974)  Coordinated activities on federal lands including wildlife management

More Recent Legislation ► National Forest Management Act (1976)  Viable populations of vertebrates must be maintained on federal lands ► Emergency Salvage Timber Act (1995)  Expedited reviews for salvage timber operations on federal lands ► Renewable Resources Extension Act (1978)  Created Cooperative Forestry Assistance Act ► Forest Conservation Relief Act (1990)  Set aside harvestable land for protection of endangered species

Early Successional Forest ► Occurs after a disturbance  Primary  Secondary ► Early successional forest wildlife  American woodcock, New England cottontail  Use fire and cutting to maintain

Forests and Fires ► Some forest trees need fire  Aspen, pitch pine, oaks ► Prescribed fire  Controls non-native plants  Creates habitat for bobwhite quail, ruffed grouse, Kirtland’s warbler ► Habitat mosaic created by fire

Forest Management ► Uneven age  Trees of different ages retained via selective cutting (size class or age classes) or thinning  Seed tree, shelterwood, clustered clearcuts ► Creates or maintains habitat for wildlife  Hooded warbler, ovenbird, eastern cottontail, fox squirrel ► Even age  Clearcutting  Promotes early successional habitat ► Size of cut influences wildlife species

Distribution of tree sizes in an even-aged forest stand (A) and an uneven-aged forest stand (B) Even- and Uneven-aged Forest Stands

Recommendations for Wildlife-Sensitive Timber Harvesting 1. Protection of important habitat features (e.g., vernal pools) 2. Enhancing vertical diversity of forests 3. Retention of forested buffers along streams 4. Retention of overstory trees 5. Retention of dead and decaying trees 6. Retention of woody debris 7. Creation of meandering edges 8. Maximizing forest interior habitat 9. Retention of old forest stands

Forest Fragmentation ► Transformation of a large expanse of forest into a number of smaller patches of smaller total area, which are isolated from one another by a matrix of habitats unlike the original. ► Creates edge--Junction of two landscape elements (plant communities, land uses, or successional stages

Edge Created by Forest Cutting

Edge Effects on Wildlife ► Birds—increased nest parasitism by brown-headed cowbirds; increased nest predation ► Small isolated forest patches may not support area-sensitive species like neotropical migrants, large mammalian predators ► Corridors or connections between patches help diminish fragmentation effects ► Forest reserves should minimize edge, be continuous, large (~ 30 ha), with minimal disturbances

Species Area Curve

Biodiversity and Forests ► Biodiversity (biological diversity)  Genetic  Species  Community/ecosystem  Landscape ► Ecosystem management  Management with biodiversity values included  Applies to variety of spatial scales (local, regional, global)

Managing Forests for Wildlife ► 5 main considerations  Biodiversity  Fragmentation  Education (forests as outdoor laboratories)  Recreation (hunting, hiking, bird watching)  Regional and Global influences (countering effects of global climate change, stabilizing soils, habitat protection)

SUMMARY Forest types, ownership, and legislation can affect the value of forests to wildlife. Plant-forest succession resulting from fire, chemicals, or management systems can have profound effects on forest wildlife. For example, uneven-aged forest management can influence wildlife distribution and abundance differently than even-aged forest management. Several practices (e.g., retention of unique habitat features or overstory trees in harvested stands) can have positive effects on forest wildlife. The impacts of forest management, however, cannot be considered solely from the stand level (e.g., stand size), but must take into consideration landscape features (e.g., characteristics of surrounding landscape). Several types of models are available to predict wildlife species occurrences and adjust forest-management regimes. Successful forest management in the future will be contingent on biodiversity conservation and ecosystem management to ensure a sustainable, productive, and healthy forest for the benefit of wildlife while simultaneously providing educational and recreational values.