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Terrestrial Biomes
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Global Air Circulation & Regional Climates
Uneven heating of the Earth’s Surface Air is more heated at the equator and less at the poles.
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Global Air Circulation & Regional Climates
Seasonal changes in temperature and precipitation
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Insolation B A C
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Solar Energy
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Seasonal shift in rainy/dry seasons
Rainy Season Seasonal shift in rainy/dry seasons
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Global Air Circulation & Regional Climates
Properties of air and water. Cell 3 South Cold, dry air falls Moist air rises — rain Cell 2 South Cool, dry air falls Cell 1 South Moist air rises, cools, and releases moisture as rain Cell 1 North Cell 2 North Cell 3 North Cold, dry Polar cap Arctic tundra 60° 30° 0° Evergreen coniferous forest Temperate deciduous forest and grassland Desert Tropical deciduous forest Equator Tropical rain forest Tropical deciduous forest
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Terrestrial Biomes of the World
Biome – a large geographical region having a defining climate to which plants show a similar physiological adaptation. Tundra Boreal Forest Temperate Forest Prairies and Grasslands Chaparral Desert Tropical Forest Tropical Mountains
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Temperature and precipitation regulate plant growth, thus the regional distribution of biomes.
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Global Distribution of Terrestrial Biomes
Global Net Primary Productivity For Terrestrial Biomes
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Net Primary Production of Terrestrial Biomes
NPP (g C/m2/yr) Tropical Rain Forest 900 Tropical Dry Forest 675 Temperate Evergreen Forest 585 Temperate Deciduous Forest 540 Boreal Forest 360 Tropical Grasslands 315 Cultivated land (USA) 290 Chaparral 270 Prairie 225 Tundra Desert 32 Extreme Desert 1.5
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Tundra Windswept expanses where nothing stands taller than grasses and sedges. Average temperature is -5°C. Water is held as ice for most of the year; growing season is from May to August Plant growth is inhibited Decomposition and nutrient cycling is very slow; soils are rich in organic matter Each year, only the top meter defrosts, below that the ground remains frozen year round - Permafrost
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Tundra The organic rich soil in the tundra called peat.
Due to the short growing season, all growth and reproduction must happen in a short period. Young shoots are thus loaded with nutrients Makes good grazing Food is so abundant at this time that wading birds, ducks, geese and swans commonly migrate to the tundra; herds of caribou and reindeer arrive and support biting flies, midges, and mosquitoes. Insects spend the winter as eggs, larvae or pupae. Birds usually feast on the abundant newly-hatched insects
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Tundra Although there are a lot of individual birds, mammals, and insects, there are not many types of species. Few plant species have evolved to withstand the harsh tundra climate. Tundra has a low species diversity (# of species) Naturally low diversity does not mean unimportant. Tundra is essential to the well-being of animals such as ducks, geese, seals, polar bears, and caribou Recovery from disturbance takes a long time Tracks from vehicles are visible for several months
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Tundra
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Boreal Forest Great fir forest of Canada and northern Eurasia.
Vegetation type is defined by seasonal expansion and contraction of the Arctic and continental polar air masses. Polar air in the winter, continental air in the summer Very cold winters, relatively warm summers so trees can survive
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Boreal Forest Many trees look like Christmas trees, not oak trees
Shape of tree is related to the ability to shed snow Broken trees are not good competitors Leaf morphology is important Needles are more durable, can photosynthesize year round conserve heat more efficiently Boreal forests are low in species diversity, but not as low as tundra
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Boreal Forest
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Temperate Forest Can be divide into evergreen and deciduous forest types. Generally have a moist cool climate. Sometimes referred to as temperate rain forests Contain giant redwoods and Douglas fir Redwoods can reach 120 m (~360 ft) in height. High-quality wood makes these forests a valuable timber resource Old Growth Forest
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Temperate Forest (Deciduous)
Hot summers, cool winters (minimum temp rarely < -12C). Rain is plentiful (75 – 200 cm / yr), long growing season, fertile soil, moderate diversity. Species include oak, beech and maple. Produce a new batch of leaves each year Not as expensive as ‘freeze-proofing’ them Early spring, a lot of light hits the forest floor and spring ephemerals take advantage of this. Soils make good farmland and supports good lumber
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Prairies and Grasslands
Climatically similar to temperate forests but are dryer. Precipitation ranges from a low 40 mm (plants seldom grow taller than 0.5m) to a high of 80 cm (plants can grow more than 2 m – tall grass prairie). Home to numerous herbivores Grazing is an important ecological factor in these areas Heat and aridity become important evolutionary factors Panting, sweating, large ears, evapotranspiration
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Prairies and Grasslands
Native grazers prevented any single plant from out competing the others. Introduced domestic grazers (cattle or sheep for example) have different grazing patterns and the floristic composition has been resorted. Soils are very rich – America’s bread basket. Fire is an important component of grasslands. Some areas may burn every three to five years (lightning) Keeps trees from becoming established Stopping these fires can alter the community, allowing trees to grow.
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Chaparral The summer drought is what drives this biomes ecology
Wet and mild winter climate, dry and mild summer Found in coastal California, Israel, parts of South America and Australia Mediterranean climate The summer drought is what drives this biomes ecology Plants either limit the water they lose, or gain as much water as possible
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Chaparral Plants can gain as much water as possible
Plants can limit water loss by producing small hard green leaves Hardness due to a waxy outer covering used to prevent water loss Most of these plants are evergreen, so they can photosynthesize during the wet part of the year By retaining leaves year round – they are able to take advantage of all rainfall Plants can gain as much water as possible Deep tap roots Extensive root system (bare ground between plants)
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Chaparral Eradication of top predators when these areas are settled leads to a proliferation of herbivores. Sometimes, native herbivores have been replaced with goats Goats are indiscriminate browsers and kill many of the plants they eat Overgrazing has reduced many Mediterranean hillsides to bare rock with very few patches of vegetation Urbanization in California
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Chaparral
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Desert Dry air descends usually at Latitudes 20 – 30 North and 20 – 30 South. Too dry to support most life forms Cloudless skies permit wide temperature fluctuations 50 C during the day, near 0 C at night. Desert plants and animals must be obsessive about water retention
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Desert Rainshadow
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Desert Desert animals usually hide from the full strength sun and come out when the temp is lower. Top predators are usually snakes and lizards – need less water than mammals Camels have a highly developed nasal structure that prevents water loss and its hair is efficient at reflecting the sun’s heat. Desert plants can not escape the sun. Evolution has shaped them to minimize exposure to the sun
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Desert
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Desert Desert plants are also designed to retain as much water as possible. Stem and leaves appear to be thick and fleshy, skin very leathery. Needles create microclimate in still air. Extensive root network; leaves are recycled
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Desert
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Tropical Forest Key feature is constant temperature
Average temperature from one month to the other is usually within ± 2C However, daytime temp may be 34 C during the day and 20 C at night. Much rain: annual rainfall ranges from 2,000 mm to 15,000 mm (79 – 591 inches; 1.6 inches per day). Wet and dry seasons Length of dry season determines the areas ecology
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Tropical Forest Long Dry Season Short Dry Season
Dry season = 6-8 months Many trees drop their leaves during the dry season not to escape cold, but to prevent overheating Short Dry Season Dry season < 3 months Tropical rain forest: 2,300 – 5,000 mm of rain per year (extreme years some places may get 15,000 mm = 50 feet).
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Tropical Forest (Rain)
Huge diversity Perhaps % of all organisms on Earth inhabit theses forests. Temperate forest may have tree species per hectare (2.47 acres); rain forests can support more than 350 – 450 tree species per hectare To fit that many tress each species may only be represented once or twice Most productive terrestrial biome High temp, moisture, uninterrupted growing season Decomposers / nutrient cycling
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Tropical Rain Forest
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Tropical Mountains Go from tropical forest to shrubs to ice as you move up in altitude. Similar phenomena seen as you move from the equator north
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Tropical Mountain
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Temperate Deciduous Forest Northern Coniferous Forest Arctic Tundra
Alpine Tundra Tropical Forest Tropical Forest Temperate Deciduous Forest Northern Coniferous Forest Arctic Tundra Low High Moisture Availability Elevation
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Tropical Mountains
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Aquatic Biomes
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Aquatic Biomes Marine Fresh Water Ocean Estuary
Rivers and Streams (Lotic) Lakes and Ponds (Lentic)
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Oceans Continental Shelf – the portion of the continental plate that lies submerged under the ocean. Usually has a gentle slope Width can vary from a few to ten kilometers Continental Slope – that area that drops from the continental shelf to the full depth of the ocean floor.
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Oceans Oceanic zone – any portion of the ocean beyond the continental shelf. Neritic zone – section of ocean that lies over the continental shelf. Littoral zone – shoreline between the high and low tide marks.
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Oceans Photic Zone – area of water that sunlight penetrates
Aphotic zone – area of water that sunlight does not penetrate
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Oceans Thermocline – vertical area where temperature abruptly changes; restricts the mixing of upper and lower water masses.
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Oceans Salinity averages 35 ppt (full strength sea water).
Due to high concentrations of sodium and chloride Ocean is more than salt and water, but most ocean waters are very poor in nutrients Phosphate, nitrate, ammonium, iron Oceans cover ~71% of Earth, but only account for 50% of the Earth’s primary production. Biological deserts not limited by water, but by nutrients Unlike terrestrial biomes, production is not higher at equator and lower at the tropics –respond to nutrient concentrations like upwellings.
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Oceans Coastal regions are much more productive than non-coastal areas. Rich nutrient input from coastal rivers Most of the worlds great fisheries come from the continental shelf Too many nutrients can lead to algal blooms, which may deoxygenate the water (eutrophication)
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Some Fish Life History Anadromous – fish that spend their adult life in salt water but spawn in freshwater Salmon, striped bass, American shad Catadromous – fish that spend their adult life in freshwater but spawn in saltwater American eel
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Sargasso Sea
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Ocean Benthic Zone Benthic zone is the ocean bottom
a thick blanket of mud that consists of fine particles that have settled from the overlying water and accumulated over millions of years. Scientists originally thought that life could not exist in the benthic zone. Too much pressure, too dark, too cold, lack of food We now know that there is a lot of life down there.
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Chemoautotrophic Organisms
Photosynthetic organisms are labeled as autotrophs, specifically phototrophs. Organisms that are able to harvest energy from inorganic compounds without photosynthesis are called chemoautotrophs. Usually sulfur-oxidizing (harvesting energy rich electrons from sulfur compounds) organisms
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Chemosynthetic Communities
Organisms use organic material seeping from the ocean floor as an energy source Colorful crab perched on top of a large tubeworm cluster at GC 354, depth 532 m. This community was first discovered on this MMS/LSU subdive, August 24, 2000
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Symbiotic Relationships
Consumer organisms found in chemosynthetic communities rely on a symbiotic relationship with chemoautotrophic bacteria. Consumer organisms must take up inorganic carbon (CO2) and sulfides and get rid of the bacterial waste. The bacteria are able to capture energy (oxidize) from the sulfides to reduce carbon dioxide (primary production). Consumers then absorb nutrients from the bacteria! Consumer organisms do not have a mouth or gut!
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Estuaries Where the river meets the sea Most productive biome on Earth
Sometimes classified freshwater, sometimes classified marine Most productive biome on Earth Support a diverse fauna including a variety of worms, oysters, crabs, and waterfowl. Serve as nursery habitat for many organisms.
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Marsh Types Fresh water marsh salinity < 1.0 ppt.
Plants are not salt tolerant and include maidencane, bulltongue, alligatorweed, cattails, and spikerush Intermediate marsh salinity averages 3.3 ppt. Plants are slightly salt tolerant and include spikerush, three-corner grass, arrowhead, cordgrass, wiregrass, roseau cane, and deer pea Brackish marsh salinity averages about 8 ppt. typically dominated by cordgrass or wiregrass Salt marsh salinity averages about 16 ppt. oyster grass is common, but few other plant species can survive
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Freshwater Marsh Brackish Marsh Intermediate Marsh Salt Marsh
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Rivers and Streams Generally represent the excess of precipitation on land areas over evaporation from them. Precipitation that falls is either evaporated, transpirated, enters the ground water supply, or flows down rivers Flow is down-hill and varies seasonally Related to rainfall and ice/snow melt Beginning of a river = the source and the end of a river = the mouth Discharge - volume of water passing a given point during a period of time Channel Width X Depth X Velocity
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Rivers and Streams Flow velocity is important in determining abiotic and biotic components. Flow related to slope and precipitation Sediment type, current strength Only certain organisms can withstand strong flow The faster the flow, the more material can be transported in the water Materials are transported by running water in three principal states Dissolved matter Suspended solids Bed load
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Stream Order Used to classify a stream in relation to tributaries, drainage area, total length, and age of water. 1 2 3 Stream Order – Strahler Method 1 1 = 2 1 2 = 2 2 2 = 3 1 3 = 3 2 3 = 3 3 3 = 4 Mississippi River is classified as a 10th or 12th order stream. Headwater stream classification matters
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Major Rivers of The World
Name Discharge 103 m3/sec Length 103 km Drainage Area 106 km2 Amazon, South America 212.40 6.44 5.78 Congo, Africa 39.65 4.70 4.01 Ganges-Brahmaputra, India 38.50 2.90 1.62 Yangtze, China 21.81 5.98 1.94 Yenisei, Russia 17.39 5.54 2.59 Mississippi North, America 17.30 6.02 3.22 Mekong, Asia 11.04 4.00 0.80 Nile, Africa 3.10 6.65 3.35
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Mississippi River (Main Stem) Atchafalaya River (Distributary)
Distributary – A smaller channel that takes water away from the main stem river. Mississippi River (Main Stem) Atchafalaya River (Distributary) Flow
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River Channel Deep Holes Sand Bars
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Oxbow Formation
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Lakes and Ponds Ponds – light can reach the entire bottom
Lakes – light can not reach some parts of the bottom
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Dissolved O2 concentration
Lake Overturn Epilimnion Hypolimnion Thermocline Summer Fall overturn 22˚ 20˚ 18˚ 8˚ 6˚ 5˚ 4˚C 0˚ 2˚ 4˚ Winter Spring overturn Dissolved O2 concentration High Medium Low
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