3. Metapopulation dynamics Rate at which subpopulations go extinct Probability of each subpopulation going extinct Proportion of patches that are occupied Rate of colonization of empty patches Probability of colonization (constant that reflects rate of dispersal between patches) Proportion of patches that are occupied

4. Metapopulation dynamics Proportion of patches that are occupied Rate of colonization of empty patches Rate at which subpopulations go extinct

7. Lotka-Volterra models of interspecific competition Population growth with carrying capacity: Competitive impact of species 2 on species 1 =  N 2  = Impact per individual of species 2 on 1 Competitive impact of species 1 on species 2 =  N 1

9. For species 1 For species 2

10. >> species 1 can’t grow >> species 2 can’t grow

18. Competition between two diatoms, Asterionella formosa and Synedra ulna for silica.