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Chapter 3: Myology
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Learning Objectives Compare and contrast the three types of muscle tissue in the human body. Discuss the five functions of skeletal muscle. Compare and contrast parallel and pennate fiber arrangements and give an example of each. Identify the six factors that make up muscle names. Give examples using each factor. Explain the contribution of each of the five properties of skeletal muscle tissue to human movement. Identify the major macroscopic and microscopic structures of muscle tissue and describe the function of each.
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Learning Objectives (cont’d)
List the events that lead to a skeletal muscle contraction and identify all chemicals necessary in the process. Discuss the factors that influence the amount of force produced by a muscle. Compare and contrast slow twitch, fast twitch, and intermediate muscles fibers. Compare and contrast isometric and isotonic muscle contractions. Compare and contrast concentric and eccentric muscles contractions. Discuss the functional interrelationships between agonist, synergist, and antagonist muscles.
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Learning Objectives (cont’d)
Identify the major skeletal muscles of the human body. Identify the components of a lever and give an example of each type of lever in the human body. Identify and describe the anatomical structures of proprioception in the human body. Define and demonstrate active, passive, and resisted range of motion. Explain the purpose of performing active, passive, and resisted range of motion.
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Types of Muscle Tissue Smooth Muscle (involuntary)
In walls of hollow organs, vessels, & respiratory passageways; functions in digestion, reproduction, circulation, & breathing Has no striations (visible alternating dark & light fibers) Cardiac Muscle (involuntary) Makes up wall of heart; creates pulsing action to circulate blood Unique in that electrical impulse travels from cell to cell Skeletal Muscle (voluntary) Connected to bones; produces movement at joints
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Types of Muscle Tissue (cont’d)
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Skeletal Muscle Functions
Motion: Primary function is to exert pull on bones, creating motion Posture: Maintain upright posture against gravity Protection: Protect underlying structures in areas where bones do not Thermogenesis: Produce body heat Vascular Pump: Help propel circulation of lymph & venous blood via contractions of skeletal muscles
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Fiber Direction and Naming Muscles
Parallel Arrangements Fibers equal in length that do not intersect Enables entire muscle to shorten equally & in same direction; maximizes range of motion Fusiform: thick central belly with tapered ends (brachialis, biceps brachii) Circular: fiber arrangements surround an opening to form a sphincter; designed to contract & close passages or relax & open them (orbicularis oris, sphincter ani) Triangular: broad at base, converging to single point; fan- shaped arrangement allows diverse actions (pectoralis major, trapezius)
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Fiber Direction and Naming Muscles (cont’d)
Pennate Arrangements Feather-shaped, with shorter fibers intersecting a central tendon Maximize number of fibers in an area, cross-sectional area, & force production Unipennate: Fibers run obliquely from one side of central tendon; look like half a feather (tibialis posterior, biceps femoris) Bipennate: Fibers run obliquely along both sides of central tendon; look like a full feather (rectus femoris) Multipennate: Multiple tendons with oblique muscle fibers on both sides; produce least amount of force (deltoid)
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Fiber Direction and Naming Muscles (cont’d)
Oblique (slanting): external oblique Rectus (straight): rectus abdominis Location Brachii (arm): biceps brachii Femoris (thigh): rectus femoris Pectoralis (chest): pectoralis major Abdominus (abdomen): rectus abdominus
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Fiber Direction and Naming Muscles (cont’d)
Action Flexor: flexor carpi radialis Extensor: extensor digitorum Pronator: pronator teres Size Major/minor: pectoralis major & minor Maximus/medius/minimus: gluteus maximus, etc. Longus/brevis/tertius: peroneus longus, etc.
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Fiber Direction and Naming Muscles (cont’d)
Shape Trapezoid: trapezius Delta: deltoid Saw-shaped: serratus anterior Number of Heads Biceps (2 heads): biceps brachii Triceps (3 heads): triceps brachii Quadriceps (4 heads): quadriceps (4 anterior thigh muscles)
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Skeletal Muscle Properties
Extensibility Ability to stretch without sustaining damage Allows muscles to lengthen when relaxed Elasticity Ability to return to original shape after lengthening or shortening Maintains a specific shape & geometry in muscles despite malleable nature Excitability Ability to respond to a stimulus by producing electrical signals Allows muscle to contract & function
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Skeletal Muscle Properties (cont’d)
Conductivity Ability to propagate electrical signals, including action potentials Allows action potential to be transmitted along muscle cell, activating tissue & initiating muscle contraction Contractility Ability to shorten & thicken, thus producing force, in response to a specific stimulus Allows force production & movement
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Anatomy of Skeletal Muscle Tissue
Macroscopic Anatomy Connective tissue wrappings support, protect, & separate portions of muscle & whole muscles Fibers: individual muscle cells Endomysium: sheath of connective tissue enveloping fibers Fascicles: bundles of grouped muscle fibers Perimysium: layer of connective tissue enveloping fascicles Epimysium: layer of deep fascia enveloping bundle of fascicles Musculotendinous junction: point where epimysium converges to form a tendon Muscle belly: portion of muscle between tendons
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Anatomy of Skeletal Muscle Tissue (cont’d)
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Anatomy of Skeletal Muscle Tissue (cont’d)
Microscopic Anatomy Sarcolemma: cell membrane; regulates chemical transport into & out of fiber Sarcoplasm: a gelatinous substance surrounding structures within fiber; cytoplasm of muscle cells Nuclei: structures within a fiber that contain functional information for cell & control its operations Myofibrils: specialized contractile proteins that make skeletal muscle tissue appear striated; two filaments: thin & thick Sarcomere: functional unit of muscle fiber; their shortening causes contraction
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Anatomy of Skeletal Muscle Tissue (cont’d)
Microscopic Anatomy Mitochondria: produce adenosine triphosphate (ATP), a compound that stores energy needed for muscle contraction Transverse tubules: network of tubules that run at right angles to sarcomeres & transmit nerve impulses from sarcolemma to cell interior Sarcoplasmic reticulum: network of fluid-filled chambers that covers each myofibril like a lacy sleeve; stores calcium ions
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Physiology of Muscle Contraction
Steps Involved in Initiating Muscle Contraction 1. A neuron sends electrical signal (action potential) down axon 2. Signal reaches ends of axon branches, stimulates release of neurotransmitter acetylcholine (ACh) 3. ACh molecules cross synaptic cleft & bind with receptors in sarcolemma 4. A muscle action potential travels along sarcolemma & down transverse tubules
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Physiology of Muscle Contraction (cont’d)
Neuromuscular junction
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Physiology of Muscle Contraction (cont’d)
Sliding Filament Theory 1. Action potential travels to sarcoplasmic reticulum & releases calcium ions into sarcoplasm 2. Calcium ions bind with troponin, moving aside tropomyosin protein strands covering binding sites on actin filament 3. Myosin heads are charged with energy from breakdown of ATP 4. Energy binds myosin heads to active receptor sites on actin filament, making connections called cross-bridges 5. Ratcheting action (power stroke) occurs as myosin heads pull sarcomere together, shortening the strand 6. Myosin heads bind more ATP, providing energy needed to release hold on actin strand; process creates contractions
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Physiology of Muscle Contraction (cont’d)
Events of muscle contraction
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Physiology of Muscle Contraction (cont’d)
Sliding filament mechanism
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Physiology of Muscle Contraction (cont’d)
Factors Affecting Force Production Motor unit recruitment Motor neuron: neuron responsible for initiating motion Motor unit: motor neuron & all fibers it controls Few fibers = fine movement, less power; many fibers = no fine movement, greater power One muscle is typically composed of multiple motor units Body can control amount of force by varying number & size of motor units recruited Summation: process of recruiting more & more motor units
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Physiology of Muscle Contraction (cont’d)
Motor unit
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Physiology of Muscle Contraction (cont’d)
Factors Affecting Force Production Cross-Sectional Area As myofibrils become larger, muscles increase in cross- sectional area & can generate more force Fiber Arrangement Pennate fiber arrangements generate more total force than parallel fiber arrangements Muscle Length Shortened muscles have decreased ability to produce force Muscles at resting length can produce greatest force
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Physiology of Muscle Contraction (cont’d)
Length-tension relationship
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Skeletal Muscle Fiber Types
Slow Twitch Fibers Contract slowly but are resistant to fatigue Rely on aerobic energy production Used for long-duration activities (walking, jogging) Fast Twitch Fibers Contract rapidly & powerfully but fatigue quickly Larger in diameter than slow twitch fibers due to having more myofilaments Rely on anaerobic energy production Used for short-duration activities (sprinting, lifting)
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Skeletal Muscle Fiber Types (cont’d)
Intermediate Fibers Have characteristics of both slow twitch & fast twitch fibers May adapt to body’s demands “Reservists” waiting to be called up when & where need arises Distribution of Fiber Types Intermingled & genetically determined Those with higher concentration of slow twitch fibers have longer, leaner muscles & predisposition to endurance activities Those with higher concentration of fast twitch fibers have larger, thicker muscles & predisposition for sprinting or body building
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Types of Muscle Contractions
Isometric Contractions Tension is generated in muscle, but muscle length & joint angle don’t change (pushing against an immovable object) Used to stabilize joints Isotonic Contractions Change muscle length & create movement Two types Concentric: muscle shortens (lifting a book off a table) Eccentric: muscle lengthens (slowly lowering book to table)
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Muscle Relationships Agonists Synergists Antagonists
Create joint movement (prime movers) Synergists Assist agonist in creating movement Stabilize, steer, or contribute to a particular joint movement Muscles that have same action or actions are considered synergists Antagonists Perform opposite actions of agonist (i.e., flexion/extension) Critical for proper posture & controlling & finishing movements
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Muscles of the Human Body
A. Anterior view.
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Muscles of the Human Body (cont’d)
B. Posterior view.
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Levers in the Human Body
A. First-class levers. B. Second-class levers. C. Third- class levers.
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Proprioception Proprioception: overall awareness of body position
Types of Proprioceptors Muscle Spindles: distributed throughout skeletal muscle & monitor changes in tissue length Golgi Tendon Organs: woven into connective tissue in tendons & monitor changes in muscle tension Vestibular Apparatus: located in inner ear & provides feedback about head position Mechanoreceptors: specialized nerve endings that deform in response to pressure & indicate position & movement of associated structures
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Range of Motion Range of motion: extent of movement possible at a joint Active Range of Motion Client moves a given body part through its possible motions independently Passive Range of Motion Client rests while therapist moves joint through its possible movements Resisted Range of Motion Client meets resistance from practitioner in attempting to produce movement at a joint
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