An Introduction to Ecology and the Biosphere Chapter 52 An Introduction to Ecology and the Biosphere

Overview: The Scope of Ecology Ecology is the scientific study of the interactions between organisms and the environment These interactions determine distribution of organisms and their abundance The Scope of Ecological Research Ecologists work at levels ranging from individual organisms to the planet

Fig. 52-2a Organismal ecology studies how an organism’s structure, physiology, and (for animals) behavior meet environmental challenges (Chapter 51) Figure 52.2 The scope of ecological research

Fig. 52-2b A population is a group of individuals of the same species living in an area Population ecology focuses on factors affecting how many individuals of a species live in an area Chapter 53 Figure 52.2 The scope of ecological research

A community is a group of populations of different species in an area Fig. 52-2c A community is a group of populations of different species in an area Community ecology deals with the whole array of interacting species in a community Chapter 54 Figure 52.2 The scope of ecological research

Fig. 52-2d An ecosystem is the community of organisms in an area and the physical factors with which they interact Ecosystem ecology emphasizes energy flow and chemical cycling among the various biotic and abiotic components Chapter 55 Figure 52.2 The scope of ecological research

Northern Coniferous Forest Fig. 52-2d Biomes are on of the worlds’ major communities classified according to the predominant vegetation and characterized bv adaptations of organisms to that particular environment Northern Coniferous Forest Figure 52.2 The scope of ecological research Coral Reef

A landscape is a mosaic of connected ecosystems Fig. 52-2e A landscape is a mosaic of connected ecosystems Landscape ecology deals with arrays of ecosystems and how they are arranged in a geographic region Figure 52.2 The scope of ecological research

The biosphere is the global ecosystem, the sum of all the planet’s ecosystems Global ecology examines the influence of energy and materials on organisms across the biosphere Figure 52.2 The scope of ecological research

Concept 52.2: Interactions between organisms and the environment limit the distribution of species Ecologists have long recognized global and regional patterns of distribution of organisms within the biosphere Biogeography is a good starting point for understanding what limits geographic distribution of species Ecologists recognize two kinds of factors that determine distribution: biotic, or living factors, and abiotic, or nonliving factors

Kangaroos/km2 0–0.1 0.1–1 1–5 5–10 10–20 > 20 Limits of Fig. 52-5 Kangaroos/km2 0–0.1 0.1–1 1–5 5–10 10–20 > 20 Limits of distribution Figure 52.5 Distribution and abundance of the red kangaroo in Australia, based on aerial surveys

Fig. 52-6 Ecologists consider multiple factors when attempting to explain the distribution of species Why is species X absent from an area? Yes Area inaccessible or insufficient time Yes Does dispersal limit its distribution? Habitat selection Yes Predation, parasitism, competition, disease Does behavior limit its distribution? Chemical factors No Do biotic factors (other species) limit its distribution? Water Oxygen Salinity pH Soil nutrients, etc. No Do abiotic factors limit its distribution? No Temperature Light Soil structure Fire Moisture, etc. Physical factors Figure 52.6 Flowchart of factors limiting geographic distribution

Dispersal contributes to global distribution of organisms Fig. 52-7 Current Dispersal is movement of individuals away from centers of high population density or from their area of origin Dispersal contributes to global distribution of organisms 1970 1966 1965 1960 1961 1958 1943 1937 1951 1956 Figure 52.7 Dispersal of the cattle egret in the Americas 1970

Behavior and Habitat Selection Some organisms do not occupy all of their potential range Species distribution may be limited by habitat selection behavior Biotic Factors Biotic factors that affect the distribution of organisms may include: Interactions with other species Predation Competition

Does feeding by sea urchins limit seaweed distribution? Fig. 52-8 Does feeding by sea urchins limit seaweed distribution? RESULTS 100 Both limpets and urchins removed 80 Sea urchin Only urchins removed 60 Seaweed cover (%) Limpet 40 Figure 52.8 Does feeding by sea urchins limit seaweed distribution? Only limpets removed 20 Control (both urchins and limpets present) August 1982 February 1983 August 1983 February 1984

Abiotic Factors Abiotic factors affecting distribution of organisms include: Temperature Water Sunlight Wind Rocks and soil Most abiotic factors vary in space and time

Temperature Environmental temperature is an important factor in distribution of organisms because of its effects on biological processes Cells may freeze and rupture below 0°C, while most proteins denature above 45°C Mammals and birds expend energy to regulate their internal temperature Water Water availability in habitats is another important factor in species distribution Desert organisms exhibit adaptations for water conservation

Salinity Salt concentration affects water balance of organisms through osmosis Few terrestrial organisms are adapted to high-salinity habitats Sunlight Light intensity and quality affect photosynthesis Water absorbs light, thus in aquatic environments most photosynthesis occurs near the surface In deserts, high light levels increase temperature and can stress plants and animals

Fig. 52-9 Organisms living at high elevations are exposed to high levels of ultraviolet radiation. They face other challenges as well, including freezing temperatures and strong winds, which increase water loss and inhibit the growth of limbs on the windward side of trees Figure 52.9 Alpine tree

Rocks and Soil Many characteristics of soil limit distribution of plants and thus the animals that feed upon them: Physical structure pH Mineral composition Climate Four major abiotic components of climate are temperature, water, sunlight, and wind The long-term prevailing weather conditions in an area constitute its climate Macroclimate consists of patterns on the global, regional, and local level Microclimate consists of very fine patterns, such as those encountered by the community of organisms underneath a fallen log

Bodies of Water The Gulf Stream carries warm water from the equator to the North Atlantic Oceans and their currents and large lakes moderate the climate of nearby terrestrial environments During the day, air rises over warm land and draws a cool breeze from the water across the land As the land cools at night, air rises over the warmer water and draws cooler air from land back over the water, which is replaced by warm air from offshore

Labrador current Gulf stream Equator water Warm Cold water Fig. 52-11 Figure 52.11 The great ocean conveyor belt Cold water

Warm air over land rises. Fig. 52-12 Air cools at high elevation. 2 Cooler air sinks over water. 3 Warm air over land rises. 1 Cool air over water moves inland, replacing rising warm air over land. 4 Figure 52.12 Moderating effects of a large body of water on climate

Concept 52.3: Aquatic biomes are diverse and dynamic systems that cover most of Earth Biomes are the major ecological associations that occupy broad geographic regions of land or water Varying combinations of biotic and abiotic factors determine the nature of biomes Aquatic biomes account for the largest part of the biosphere in terms of area They can contain fresh water or salt water (marine) Oceans cover about 75% of Earth’s surface and have an enormous impact on the biosphere

Tropic of Cancer Equator Tropic of Capricorn Fig. 52-15 Lakes Coral reefs Rivers Oceanic pelagic and benthic zones Estuaries 30ºN Intertidal zones Tropic of Cancer Equator Tropic of Capricorn 30ºS Figure 52.15 The distribution of major aquatic biomes

Stratification of Aquatic Biomes Fig. 52-16 Stratification of Aquatic Biomes Many aquatic biomes are stratified into zones or layers defined by light penetration, temperature, and depth Intertidal zone Neritic zone Oceanic zone Littoral zone Limnetic zone Photic zone 200 m Continental shelf Pelagic zone Benthic zone Aphotic zone Photic zone Pelagic zone Benthic zone Aphotic zone Figure 52.16 Zonation in aquatic environments 2,000–6,000 m Abyssal zone (a) Zonation in a lake (b) Marine zonation

The upper photic zone has sufficient light for photosynthesis while the lower aphotic zone receives little light The organic and inorganic sediment at the bottom of all aquatic zones is called the benthic zone The communities of organisms in the benthic zone are collectively called the benthos Detritus, dead organic matter, falls from the productive surface water and is an important source of food The most extensive part of the ocean is the abyssal zone with a depth of 2,000 to 6,000 m

Aquatic Biomes Major aquatic biomes can be characterized by their physical environment, chemical environment, geological features, photosynthetic organisms, and heterotrophs

Oligotrophic lakes are nutrient-poor and generally oxygen-rich Eutrophic lakes are nutrient-rich and often depleted of oxygen if ice covered in winter Figure 52.18 Aquatic biomes An oligotrophic lake in Grand Teton National Park, Wyoming A eutrophic lake in the Okavango Delta, Botswana

Okefenokee National Wetland Reserve in Georgia Wetlands A wetland is a habitat that is inundated by water at least some of the time and that supports plants adapted to water-saturated soil Wetlands can develop in shallow basins, along flooded river banks, or on the coasts of large lakes and seas Wetlands are among the most productive biomes on earth and are home to diverse invertebrates and birds!!!!!!! Figure 52.18 Aquatic biomes Okefenokee National Wetland Reserve in Georgia

A fast current causes an increase in oxygen levels!!!! Fig. 52-18d Streams and Rivers The most prominent physical characteristic of streams and rivers is current A fast current causes an increase in oxygen levels!!!! Figure 52.18 Aquatic biomes A headwater stream in the Great Smoky Mountains The Mississippi River far from its headwaters

An estuary is a transition area between river and sea Fig. 52-18f Estuaries An estuary is a transition area between river and sea Salinity varies with the rise and fall of the tides Estuaries are nutrient rich and highly productive An abundant supply of food attracts marine invertebrates and fish Figure 52.18 Aquatic biomes An estuary in a low coastal plain of Georgia

Fig. 52-19 Concept 52.4: The structure and distribution of terrestrial biomes are controlled by climate and disturbance Tropical forest Savanna Desert Chaparral 30ºN Tropic of Cancer Temperate grassland Equator Temperate broadleaf forest Tropic of Capricorn Northern coniferous forest 30ºS Tundra Figure 52.19 The distribution of major terrestrial biomes High mountains Polar ice Climate is very important in determining why terrestrial biomes are found in certain areas Biome patterns can be modified by disturbance such as a storm, fire, or human activity

General Features of Terrestrial Biomes and the Role of Disturbance Terrestrial biomes are often named for major physical or climatic factors and for vegetation Terrestrial biomes can be characterized by distribution, precipitation, temperature, plants, and animals Terrestrial biomes usually grade into each other, without sharp boundaries Vertical layering is an important feature of terrestrial biomes, and in a forest it might consist of an upper canopy, low-tree layer, shrub understory, ground layer of herbaceous plants, forest floor, and root layer Layering of vegetation in all biomes provides diverse habitats for animals Biomes are dynamic and usually exhibit extensive patchiness

Tropical Forest In tropical rain forests, rainfall is relatively constant, while in tropical dry forests precipitation is highly seasonal Tropical forests are vertically layered and competition for light is intense Tropical forests are home to millions of animal species, including an estimated 5–30 million still undescribed species of insects, spiders, and other arthropods. (Highest biodiversity of terrestrial biomes) Soil is very poor in nutrients because nutrients are “tied up” in vegetation

A tropical rain forest in Borneo Fig. 52-21a Figure 52.21 Terrestrial biomes For the Discovery Video Trees, go to Animation and Video Files. A tropical rain forest in Borneo

Desert Precipitation is low and highly variable, generally less than 30 cm per year; deserts may be hot or cold Desert plants are adapted for heat and desiccation tolerance, water storage, and reduced leaf surface area Common desert animals include many kinds of snakes and lizards, scorpions, ants, beetles, migratory and resident birds, and seed-eating rodents; many are nocturnal

A desert in the southwestern United States Fig. 52-21b Figure 52.21 Terrestrial biomes A desert in the southwestern United States

Savanna (have not seen on ap) Savanna precipitation and temperature are seasonal Grasses and forbs make up most of the ground cover Common inhabitants include insects and mammals such as wildebeests, zebras, lions, and hyenas Chaparral (have not seen on ap) Chaparral climate is highly seasonal, with cool and rainy winters and hot dry summers The chaparral is dominated by shrubs, small trees, grasses, and herbs; many plants are adapted to fire and drought Animals include amphibians, birds and other reptiles, insects, small mammals and browsing mammals

Fig. 52-21c Figure 52.21 Terrestrial biomes A savanna in Kenya

An area of chaparral in California Fig. 52-21d Figure 52.21 Terrestrial biomes An area of chaparral in California

Temperate Grassland (have not seen on ap) Temperate grasslands are found on many continents (key word is temperate) Winters are cold and dry, while summers are wet and hot The dominant plants, grasses and forbs, are adapted to droughts and fire Native mammals include large grazers and small burrowers

Sheyenne National Grassland in North Dakota Fig. 52-21e Sheyenne National Grassland in North Dakota Figure 52.21 Terrestrial biomes

Northern Coniferous Forest (also known as Taiga or Boreal Forest) The northern coniferous forest, or taiga, extends across northern North America and Eurasia and is the largest terrestrial biome on Earth Winters are cold and long while summers may be hot The conical shape of conifers prevents too much snow from accumulating and breaking their branches Animals include migratory and resident birds, and large mammals

Rocky Mountain National Park in Colorado Fig. 52-21f Figure 52.21 Terrestrial biomes Rocky Mountain National Park in Colorado

Temperate Broadleaf Forest (have not seen on ap) Winters are cool, while summers are hot and humid; significant precipitation falls year round as rain and snow A mature temperate broadleaf forest has vertical layers dominated by deciduous trees in the Northern Hemisphere and evergreen eucalyptus in Australia Mammals, birds, and insects make use of all vertical layers in the forest In the Northern Hemisphere, many mammals hibernate in the winter

National Park in North Carolina Fig. 52-21g Great Smoky Mountains National Park in North Carolina Figure 52.21 Terrestrial biomes

Tundra Tundra covers expansive areas of the Arctic; alpine tundra exists on high mountaintops at all latitudes Winters are long and cold while summers are relatively cool; precipitation varies Permafrost, a permanently frozen layer of soil, prevents water infiltration Vegetation is herbaceous (mosses, grasses, forbs, dwarf shrubs and trees, and lichen) and supports birds, grazers, and their predators

Denali National Park, Alaska, in autumn Fig. 52-21h Figure 52.21 Terrestrial biomes Denali National Park, Alaska, in autumn

Fig. 52-T1

Fig. 52-UN3

100 Mean height (cm) 50 3,000 Altitude (m) 2,000 Sierra Nevada Fig. 52-UN2 100 Mean height (cm) 50 3,000 Altitude (m) 2,000 Sierra Nevada 1,000 Great Basin Plateau Seed collection sites

You should now be able to: Distinguish among the following types of ecology: organismal, population, community, ecosystem, and landscape Explain how dispersal may contribute to a species’ distribution Distinguish between the following pair of terms: biotic and abiotic factors Explain how a body of water might affect regional climatic conditions Define the following terms: photic zone, aphotic zone, benthic zone, abyssal zone, List and describe the characteristics of the major aquatic biomes List and describe the characteristics of the major terrestrial biomes Compare the vertical layering of a forest and grassland