Suborder Gobiodei: The Gobies Thomas J Bland Olatunde Akinwale 21 November, 2008 www.eol.org
Gobiodei 9 families, 270 genera, 2211 species Benthic Mostly Marine Small bodied
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Gobiodei Swim bladder absent Pelvic fins below pectoral fins Fused (usually) Lateral line reduced Cephalic canals Can be highly variable among species
Phylogenetics Gobioidei is monophyletic Families are not Thacker 2003
Loach Gobies Family Rhyacicthyidae 1 genus, 2 species Most primitive goby Retains well developed lateral line www.fishbase.org www.eol.org
Freshwater Sleepers Family Odontobutidae 5 genera, 15 species May not be monophyletic www.fishbase.org www.eol.org
Sleepers Family Eleotridae 35 genera, 155 species www.nzfreshwater.org Family Eleotridae 35 genera, 155 species Marine, brackish, and freshwater Pelvic fins separate Oxyeleotris marmorata reaches 66cm www.eol.org
Sand Gobies Family Kraemeriidae 2 genera, 8 species Elongate body Burrow into sand animaldiversity.ummz.umich.edu www.eol.org
Gobies, True Gobies Family Gobiidae 210 genera, >2000 species 10-20 new species/year Round goby (Neogobius melanostomus) animaldiversity.ummz.umich.edu www.eol.org
Wormfishes Family Microdesmidae 5 genera, 20 species Very elongate Burrow into substrate www.fishbase.org www.eol.org
Infantfishes Family Schindleriidae 1 genus, 3 species Highly Paedomorphic Transparent, some bones/cartilage does not develop Lightest, shortest lived vertebrate www.fishbase.org www.eol.org
Round Goby (Neogobius melanostomus) www.eol.org
Round Goby (Neogobius melanostomus) Ballast water introduction www.eol.org
Round Goby High reproductive rate Aggressive Displaces native benthic fishes Eat fish eggs and native mussels Native predator of Zebra Mussel (Driessena polymorpha) animaldiversity.ummz.umich.edu
Shrimp Goby
Goby vids http://video.google.com/videoplay?docid=1188610441663749549&ei=lrgkScPyJoGE_AHqrrDxBg&q=shrimp+goby&hl=en http://www.youtube.com/watch?v=taChNbtKIV0 Forward the second video to 0:30
Mudskippers Subfamily Oxudercinae 10 genera, 39 species Distribution ranges from Tropical and subtropical Intertidal soft-bottomed Shores of West Africa and Indo-Pacific region.
Mudskippers (cont’d) Some species possess adaptations to terrestrial environment and amphibious life. In particular, periophthalms (genus Periophthalmus) and boleophthalms (genus Boleophthalmus) include species that inspired popular and scientific imagination since the dawn of the Natural Sciences.
Mudskippers (cont’d) Periophthalmus argentilineatus Boleophthalmus pectinirostris http://www.youtube.com/watch?v=9S4TA9RfDb8&feature=related
Questions What adaptations do gobies have for a benthic lifestyle? Why are loach gobies considered the most primitive gobies? Why is Gobiidae so speciose? Why do experts think there may be lots of undiscovered species in Gobiidae? Where is the round goby native to? What role does the goby play in the shrimp goby/shrimp mutualism?