Evolution and Darwin

Evolution earliest formsdiversityWhat has it done? Transformed life on earth from it’s earliest forms to the vast diversity that characterizes it today. changegenes !!!!!!!!DEFINITION: A gradual change in the genes of a species over time!!!!!!!!

Old Theories of Evolution Jean-Baptiste LamarckJean-Baptiste Lamarck 1809 “The inheritance of acquired characteristics”“The inheritance of acquired characteristics” (Said acquired changes were passed to offspring) Means changes are adaptations to environment acquired during animal’s lifetime

“Law of Use and Disuse” develop characteristicspasses offspringUsing or not using its body parts, an individual tends to develop certain characteristics, which it passes on to its offspring. Example: A giraffe acquired its long neck because its ancestor stretched higher and higher into the trees to reach leaves, and that the animal’s increasingly lengthened neck was passed on to its offspring.

Mistakes of Lamarck Can you think of what’s wrong with his theory?

Charles Darwin a.Influenced by Charles Lyell “Principles of Geology”. a.Influenced by Charles Lyell who published “Principles of Geology”. DarwinThis publication led Darwin to realize that natural forces gradually change Earth’s surface and that the forces of the past are still operating in modern times.

Charles Darwin b. Born Feb. 12, 1809 c.Joined Crew of HMS Beagle, for Voyage around world Naturalist Avid Collector of Plants & Animals Astounded By Variety of Life

HMS Beagle A reconstruction of the HMS Beagle sailing off Patagonia

HMS Beagle Route Darwin left England in 1831 and returned 1836

The Galapagos Islands Small Group of Volcanic Islands 1000 km NW of South America Very Different Climates & Geography from Mainland Animals On Islands Unique Tortoises Iguanas Finches

Finches on Galapagos Islands Island Finches looked like mainland finch More Types of Finches appeared on the islands where the available food was different (seeds, nuts, berries, insects…) Finches had different types of beaks adapted to their type of food gathering 14 Different Types of Finches

Charles Darwin Wrote in 1859“On the Origin of Species by Means of Natural Selection”Wrote in 1859:“On the Origin of Species by Means of Natural Selection” Two main points:Two main points: 1.Species were not created in their present form, but evolved from ancestral species. 2.Proposed a method (way) for evolution: NATURAL SELECTION

Natural Selection Individualsfavorabletraits environmentIndividuals with favorable traits are more likely to survive and reproduce to leave more offspring better suited for their environment. “Survival of the Fittest”Also known as “Survival of the Fittest” That organisms descend from a common ancestor but evolve (or change) to form new species “Descent with Modification”“Descent with Modification” Example:Example: English peppered moth (Biston betularia) Biston betulariaBiston betularia - light and dark phases

Mechanisms of Natural Selection How it occurs? Overproduction –All organisms have the ability to produce more offspring than possibly can survive. Ex: female fish or frog lays million of eggs. Variations – Individuals vary or differ in their traits. They may differ in size, coloration, running speed, resistance to disease, and many other traits. Competition –Resources (Food, Water, Shelter) often in short supply –Predators –Non-living environment (Temperature, Weather, Habitat, Terrain) “Struggle for Existence” Only the Strongest will survive. “Survival of the Fittest”

The Peppered Moth 1.What is the Industrial Revolution? 2.Before the Industrial Revolution, what did the majority of moths look like? 3.Why were the less populated moths so few in numbers? 4.What happened to the trees during the Industrial Revolution? 5.What happened to the moth population after the Industrial Revolution? 6.Why did this change occur?

Artificial Selection selective breedingThe selective breeding of domesticated plants and animals by man. Question:Question: What’s the ancestor of the domesticated dog? Answer:WOLFAnswer: WOLF

Evidence of Evolution 1. Biogeography: Geographical distribution of species. 2. Fossil Record: Fossils and the order in which they appear in layers of sedimentary rock (strongest evidence).

Eastern Long Necked Turtle

Evidence of Evolution 3. Taxonomy: Classification of life forms. 4. Homologous structures: Structures that are similar because of common ancestry (comparative anatomy)

Evidence of Evolution 5. Comparative embryology: Study of structures that appear during embryonic development. 6. Molecular biology: DNA and proteins (amino acids)

Population Genetics sciencegenetic changeThe science of genetic change in population. Remember:Remember:Hardy-Weinberg equation.

Population individuals same speciesA localized group of individuals belonging to the same species.

Species populationsindividuals interbreed viableA group of populations whose individuals have the potential to interbreed and produce viable offspring.

Gene Pool collection of genesThe total collection of genes in a population at any one time.

Hardy-Weinberg Principle conceptshuffling of genes cannot changeThe concept that the shuffling of genes that occur during sexual reproduction, by itself, cannot change the overall genetic makeup of a population.

Hardy-Weinberg Principle principle fiveThis principle will be maintained in nature only if all five of the following conditions are met: 1.Very large population 2.Isolation from other populations 3.No net mutations 4.Random mating 5.No natural selection

Hardy-Weinberg Principle Remember:Remember: equilibrium If these conditions are met, the population is at equilibrium. “No Change” or “No Evolution”.This means “No Change” or “No Evolution”.

Macroevolution higher than the species levelThe origin of taxonomic groups higher than the species level.

Microevolution population’s gene poolA change in a population’s gene pool over a secession of generations. Evolutionary changes geological timeEvolutionary changes in species over relatively brief periods of geological time.

Five Mechanisms of Microevolution 1. Genetic drift: Change in the gene pool of a small population due to chance. Two examples:Two examples: a. Bottleneck effect b. Founder effect

a. Bottleneck Effect Genetic drift disasterreduces population sizeGenetic drift (reduction of alleles in a population) resulting from a disaster that drastically reduces population size. Examples:Examples: 1.Earthquakes 2.Volcano’s

b. Founder Effect Genetic driftcolonizationGenetic drift resulting from the colonization of a new location by a small number of individuals. random changeResults in random change of the gene pool. Example:Example: 1.Islands (first Darwin finch)

Five Mechanisms of Microevolution 2. Gene Flow: Tgain or loss of alleles movement The gain or loss of alleles from a population by the movement of individuals or gametes. Immigration or emigrationImmigration or emigration.

Five Mechanisms of Microevolution 3. Mutation: Change in an organism’s DNA that creates a new allele. 4. Non-random mating: The selection of mates other than by chance. 5. Natural selection: Differential reproduction.

Modes of Action Natural selectionthree modesNatural selection has three modes of action: 1.Stabilizing selection 2.Directional selection 3.Diversifying selection Number of Individuals Size of individuals Small Large

1.Stabilizing Selection Actsextremesfavors intermediateActs upon extremes and favors the intermediate. Number of Individuals Size of individuals Small Large

2.Directional Selection Favorsone extremeFavors variants of one extreme. Number of Individuals Size of individuals Small Large

3.Diversifying Selection Favorsopposite extremesFavors variants of opposite extremes. Number of Individuals Size of individuals Small Large

Speciation evolutionThe evolution of new species.

Reproductive Barriers mechanismimpedes fertile and/or viable hybrid offspringAny mechanism that impedes two species from producing fertile and/or viable hybrid offspring. Two barriers:Two barriers: 1.Pre-zygotic barriers 2.Post-zygotic barriers

1.Pre-zygotic Barriers a. Temporal isolation: Breeding occurs at different times for different species. b. Habitat isolation: Species breed in different habitats. c. Behavioral isolation: Little or no sexual attraction between species.

1.Pre-zygotic Barriers d. Mechanical isolation: Structural differences prevent gamete exchange. e. Gametic isolation: Gametes die before uniting with gametes of other species, or gametes fail to unite.

2.Post-zygotic Barriers a. Hybrid inviability: Hybrid zygotes fail to develop or fail to reach sexual maturity. b. Hybrid sterility: Hybrid fails to produce functional gametes. c. Hybrid breakdown: Offspring of hybrids are weak or infertile.

Allopatric Speciation ancestral separatedgeographical barrier.Induced when the ancestral population becomes separated by a geographical barrier. Example:Example: Grand Canyon and ground squirrels

Adaptive Radiation Emergence of numerous species common ancestorEmergence of numerous species from a common ancestor introduced to new and diverse environments. Example:Example: Darwin’s Finches

Sympatric Speciation reproductively isolated sub- populationResult of a radical change in the genome that produces a reproductively isolated sub- population within the parent population (rare). Example: Plant evolution - polyploidExample: Plant evolution - polyploid chromosome # A species doubles it’s chromosome # to become tetraploid. reproductive sub-population Parent population

Interpretations of Speciation Two theories:Two theories: 1.Gradualist Model (Neo-Darwinian): Slow changes in species overtime. 2.Punctuated Equilibrium: Evolution occurs in spurts of relatively rapid change.

Convergent Evolution Speciesevolutionary branches very similar environments.Species from different evolutionary branches may come to resemble one another if they live in very similar environments. Example:Example: 1.Ostrich (Africa) and Emu (Australia). 2.Sidewinder (Mojave Desert) and Horned Viper (Middle East Desert)

Coevolution Evolutionary change selective forcesecond firstEvolutionary change, in which one species act as a selective force on a second species, inducing adaptations that in turn act as selective force on the first species. Example:Example: 1.Acacia ants and acacia trees 2.Humming birds and plants with flowers with long tubes

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