4I. Concept 25.1: The Origin of Life A. Introduction 1. Conditions at this time on Earth were such that life could have arisen by “spontaneous generation” or abiogenesis (life developing from nonliving materials) 2. Louis Pasteur disproved spontaneous generation in Today we accept the theory of “biogenesis” (life comes only from preexisting life). 4. It is possible that chemical and physical processes on early Earth produced the first cells. B. Earth and other planets in the solar system formed about 4.6 billion years ago.
6C. Current theory about how life arose consists of four main stages: 1 C. Current theory about how life arose consists of four main stages: 1. Abiotic synthesis of small organic molecules (monomers) 2. Joining of monomers to form macromolecules 3. Packaging of macromolecules into “protobionts,” droplets with membranes that maintained a distinct internal chemistry 4. Origin of self-replicating molecules that eventually made inheritance possible.
7D. Closer Look at these Four Main Stages 1. Abiotic synthesis of small organic molecules (monomers)In the 1920’s, A. I Oparin (Russian) and J. B. S. Haldane (British) independently postulated that the conditions on early Earth (reducing environment) favored the synthesis of organic compounds from simple molecules.Energy possibly from lightning and UV radiationIn 1953, Stanley Miller and Harold Urey tested the Oparin-Haldane hypothesis
8They conducted lab experiments that showed that the abiotic synthesis of organic molecules in a reducing atmosphere is possibleAtmosphere of primitive Earth:water vaporhydrogen gasmethaneammoniaUsed electrical sparks as source of energy
102. Joining of monomers to form macromolecules Monomers could link to form macromolecules without enzymes and other cellular equipment.Macromolecules, including polypeptides, have been produced in the lab by dripping solutions of monomers onto hot sand, clay, or rock.These conditions are thought to be similar to early Earth at deep-sea vents or when dilute solutions of monomers splashed onto fresh lava.
113. Packaging of macromolecules into “protobionts,” droplets with membranes that maintained a distinct internal chemistryTwo properties that defined life: accurate replication and metabolismProtobionts—aggregates of abiotically produced molecules surrounded by a membrane or membrane-like structure--they exhibit simple reproduction and metabolism and maintain an internal chemical environmentCan be formed spontaneously from abiotically produced organic compounds (liposomes)
134. Origin of self-replicating molecules that eventually made inheritance possible. First genetic material was probably RNA, not DNARNA molecules called ribozymes can catalyze many different reactionsHas been demonstrated that RNA sequences can evolve under abiotic condition
15II. Concept 25.2: Fossil Record of Life on Earth A. Sedimentary rocks are deposited into layers called strata and are the riches source of fossils.B. Methods used to date fossils and rocks:1. Radiometric datingCan determine absolute ages of fossilsInvolves half-life of isotopes (time required for half the parent isotope to decay)2. Radiocarbon dating (carbon-14)Used to date fossils up to 75, 000 years old
16III. Concept 25.3: Major Lineages of Life A. Oldest know fossils are stromatolites, rock-like structures composed of many layers of bacteria and sedimentDate back 3.5 billion years agoB. Prokaryotes were Earth’s sole inhabitants from 3.5 to about 2.1 billion years agoC. About 2.7 billion years ago, O2 (probably from bacteria similar to modern cyanobacteria) began accumulating in the atmosphereMany prokaryotes were doomed by the increase in atmospheric oxygen
18D. Oldest fossils of eukaryotic cells date back to 2.1 billion years E. First Eukaryotes1. EndosymbiosisProposes that mitochondria and plastids (chloroplasts and related organelles) were formerly small prokaryotes living within larger host cellsEndosymbiont—a cell that lives within a host cellProkaryotic ancestors of mitochondria and plastids probably gained entry to the host cell as undigested prey or internal parasitesEventually host and endosymbiots would have become a single organism
19Serial endosymbiosis supposes that mitochondria evolved before plastids through a sequence of endosymbiotic events
202. Evidence to support an endosymbiotic origin. of 2. Evidence to support an endosymbiotic origin of mitochondria and plastids:a. Similarities in inner membrane structures and functionsb. Division is similar in those organelles and some prokaryotesc. Their ribosomes are more similar to prokaryotic than eukaryotic ribosomesd. Circular DNA3. Earliest Multicellular EukaryotesOldest known fossils of multicellular eukaryotes are of small algae that lived about 1.2 billion years ago
21IV. Concept 25.4: Continental Drift A. Defined as the process of the Earth’s continents moving slowly over the underlying hot mantleB. Interactions between plates cause the formation of mountains and islands, and earthquakesC. Pangaea—supercontinentBreak-up lead to allopatric speciationCurrent distribution of fossils reflects the movement of continental drift-Ex: the similarity of fossils in parts of South America and Africa is consistent with the idea that these continents were formerly attached
24V. Concept 25.5: Evolutionary Effects of Development Genes A. Genes that program development control the rate, timing, and spatial pattern of changes in an organism’s form as it develops into an adultB. Changes in Rate and Timing1. HeterochronyDefined as an evolutionary change in the rate or timing of developmental eventsCan have a significant impact on body shapeContrasting shapes of human and chimpanzee skulls are the result of small changes in relative growth rates
27Can alter the timing of reproductive development relative to the development of nonreproductive organsIn paedomorphosis, the rate of reproductive development accelerates compared with somatic development-Sexually mature species may retain body features that were juvenile structures in an ancestral species
29C. Changes in Spatial Pattern Substantial evolutionary change can result from alterations in genes that control the placement and organization of body partsHomeotic genes determine such basic features as where wings and leg will develop on a bird or how a flower’s parts are arrangedHox genes are a class of homeotic genes that provide positional information during development-If expressed in the wrong location, body parts can be produced in the wrong location
30VI. Concept 25.6: Evolution is Not Goal Oriented (not on exam) A. Evolutionary NoveltiesEvolve in many stages from previously existing structuresComplex eyes evolved from simple photosensitive cells independently many timesExaptations are structures that evolve in one context but become co-opted for a different functionNatural selection can only improve a structure in the context of its current utility
31You should now be able to: Define radiometric dating, serial endosymbiosis, Pangaea, snowball Earth, exaptation, heterochrony, and paedomorphosisDescribe the contributions made by Oparin, Haldane, Miller, and Urey toward understanding the origin of organic moleculesList and discuss the 4 main stages of how life aroseExplain why RNA, not DNA, was likely the first genetic material
32Describe and suggest evidence for the major events in the history of life on Earth from Earth’s origin to 2 billion years agoExplain the function of Hox genes