1. Chapter 35: Transport in Plants CHAPTER 35 Transport in Plants

2. Chapter 35: Transport in Plants Uptake and Transport of Water and Minerals Uptake and Transport of Water and Minerals Transport of Water and Minerals in the Xylem Transport of Water and Minerals in the Xylem Transpiration and the Stomata Transpiration and the Stomata Translocation of Substances in the Phloem Translocation of Substances in the Phloem

3. Chapter 35: Transport in Plants Uptake and Transport of Water and Minerals Plant roots take up water and minerals from the soil.Plant roots take up water and minerals from the soil.3

4. Chapter 35: Transport in Plants Uptake and Transport of Water and Minerals Water moves through biological membranes by osmosis, toward cells with more negative water potential.Water moves through biological membranes by osmosis, toward cells with more negative water potential. The water potential of a cell or solution is the sum of the solute and the pressure potentials.The water potential of a cell or solution is the sum of the solute and the pressure potentials. All three parameters are expressed in megapascals (MPa). Review Figure 35.1All three parameters are expressed in megapascals (MPa). Review Figure 35.135.1 4

5. Chapter 35: Transport in Plants Figure 35.1 figure 35-01.jpg

6. Chapter 35: Transport in Plants Uptake and Transport of Water and Minerals Mineral uptake requires transport proteins.Mineral uptake requires transport proteins. Some minerals enter the plant by facilitated diffusion; others by active transport.Some minerals enter the plant by facilitated diffusion; others by active transport. A proton pump facilitates active transport of many solutes across membranes.A proton pump facilitates active transport of many solutes across membranes. Review Figure 35.2 35.2 6

7. Chapter 35: Transport in Plants Figure 35.2 figure 35-02.jpg

8. Chapter 35: Transport in Plants Uptake and Transport of Water and Minerals Water and minerals pass from the soil to the xylem via the apoplast and symplast.Water and minerals pass from the soil to the xylem via the apoplast and symplast. In the root, water and minerals may pass from the cortex into the stele only via the symplast because Casparian strips in the endodermis block movement in the apoplast.In the root, water and minerals may pass from the cortex into the stele only via the symplast because Casparian strips in the endodermis block movement in the apoplast. Review Figures 35.3, 35.4 Review Figures 35.3, 35.435.335.435.335.48

9. Chapter 35: Transport in Plants Figure 35.3 figure 35-03.jpg

10. Chapter 35: Transport in Plants Figure 35.4 figure 35-04.jpg

11. Chapter 35: Transport in Plants Transport of Water and Minerals in the Xylem Early experiments established that sap does not move via pumping action of living cells.Early experiments established that sap does not move via pumping action of living cells.11

12. Chapter 35: Transport in Plants Transport of Water and Minerals in the Xylem Root pressure is responsible for guttation and oozing of sap from cut stumps, but cannot account for the ascent of sap in trees.Root pressure is responsible for guttation and oozing of sap from cut stumps, but cannot account for the ascent of sap in trees.12

13. Chapter 35: Transport in Plants Transport of Water and Minerals in the Xylem Xylem transport is the result of combined effects of transpiration, cohesion, and tension.Xylem transport is the result of combined effects of transpiration, cohesion, and tension. Evaporation in the leaf produces tension in the surface film of water on the moist- walled mesophyll cells, thus pulling water and dissolved minerals up the xylem from the root.Evaporation in the leaf produces tension in the surface film of water on the moist- walled mesophyll cells, thus pulling water and dissolved minerals up the xylem from the root. Review Figure 35.7 35.7 13

14. Chapter 35: Transport in Plants Figure 35.7 figure 35-07.jpg

15. Chapter 35: Transport in Plants Transport of Water and Minerals in the Xylem Support for the transpiration–cohesion– tension model came from studies using a pressure bomb.Support for the transpiration–cohesion– tension model came from studies using a pressure bomb. Review Figure 35.8 35.8 15

16. Chapter 35: Transport in Plants Figure 35.8 figure 35-08.jpg

17. Chapter 35: Transport in Plants Transpiration and the Stomata Water evaporation cools leaves, but a plant cannot afford to lose too much water.Water evaporation cools leaves, but a plant cannot afford to lose too much water. Transpirational water loss is minimized by the leaves’ waxy cuticles.Transpirational water loss is minimized by the leaves’ waxy cuticles.17

18. Chapter 35: Transport in Plants Transpiration and the Stomata Stomata allow compromise between water retention and carbon dioxide uptake.Stomata allow compromise between water retention and carbon dioxide uptake. Guard cells controls size of stomatal opening.Guard cells controls size of stomatal opening. A proton pump, activated by blue light, pumps protons from guard to epidermal cells.A proton pump, activated by blue light, pumps protons from guard to epidermal cells. Guard cells take up potassium ions, causing water to follow osmotically, swelling the cells and opening the stomata.Guard cells take up potassium ions, causing water to follow osmotically, swelling the cells and opening the stomata. Carbon dioxide level and water availability also affect stomatal opening.Carbon dioxide level and water availability also affect stomatal opening. Review Figures 35.9, 35.10 35.935.1035.935.1018

19. Chapter 35: Transport in Plants Figur e 35.9 figure 35-09.jpg

20. Chapter 35: Transport in Plants Figure 35.10 figure 35-10.jpg

21. Chapter 35: Transport in Plants Transpiration and the Stomata In most plants the stomata are open during the day and closed at night.In most plants the stomata are open during the day and closed at night. CAM plants have an inverted stomatal cycle, enabling them to conserve water.CAM plants have an inverted stomatal cycle, enabling them to conserve water. Review Figure 35.11 35.11 21

22. Chapter 35: Transport in Plants Figure 35.11 figure 35-11.jpg

23. Chapter 35: Transport in Plants Translocation of Substances in the Phloem Products of photosynthesis, and some minerals, are translocated through sieve tubes in the phloem via living sieve tube members.Products of photosynthesis, and some minerals, are translocated through sieve tubes in the phloem via living sieve tube members. Translocation proceeds in both directions in the stem, although in a single sieve tube it goes only one way.Translocation proceeds in both directions in the stem, although in a single sieve tube it goes only one way. Translocation requires a supply of ATP.Translocation requires a supply of ATP.23

24. Chapter 35: Transport in Plants Translocation of Substances in the Phloem Translocation in the phloem proceeds in accordance with the pressure flow model:Translocation in the phloem proceeds in accordance with the pressure flow model: Difference in solute concentration between sources and sinks allows a difference in pressure potential along sieve tubes, resulting in bulk flow.Difference in solute concentration between sources and sinks allows a difference in pressure potential along sieve tubes, resulting in bulk flow. Review Figure 35.14, Table 35.1 35.1435.135.1435.124

25. Chapter 35: Transport in Plants Figure 35.14 – Part 1 figure 35-14a.jpg

26. Chapter 35: Transport in Plants Figure 35.14 – Part 2 figure 35-14b.jpg

27. Chapter 35: Transport in Plants Table 35.1 table 35-01.jpg

28. Chapter 35: Transport in Plants Translocation of Substances in the Phloem The pressure flow model succeeds because sieve plates are normally open, allowing bulk flow, and because neighboring cells load organic solutes into sieve tube members in source regions unloading them in sink regions.The pressure flow model succeeds because sieve plates are normally open, allowing bulk flow, and because neighboring cells load organic solutes into sieve tube members in source regions unloading them in sink regions. Review Figure 35.15 35.15 28

29. Chapter 35: Transport in Plants Figure 35.15 – Part 1 figure 35-15a.jpg

30. Chapter 35: Transport in Plants Figure 35.15 – Part 2 figure 35-15b.jpg

31. Chapter 35: Transport in Plants Translocation of Substances in the Phloem The distribution and properties of plasmodesmata differ between source and sink tissues.The distribution and properties of plasmodesmata differ between source and sink tissues. It may become possible to regulate plasmodesmata in crop plants.It may become possible to regulate plasmodesmata in crop plants.31