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Transport In Plants
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Water Potential The physical property predicting the direction in which water will flow ____________________ Pressure water moves from high water potential to low water potential Solute Concentration
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Water Potential (a) Left Side Right Side Water moves to the right
Pure Water = 0 Water Potential Right Side Negative Water Potential 0 pressure - solute (has solutes) Water moves to the right
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Water Potential (b) Left Side Right Side Water is at equilibrium
Pure Water = 0 Water Potential Right Side 0 Water Potential + pressure equal to solute conc. - solute (has solutes) Water is at equilibrium
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Water Potential (c) Left Side Right Side Water moves to the left
Pure Water = 0 Water Potential Right Side Positive Water Potential + pressure more than solute conc. - solute (has solutes) Water moves to the left
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Water Potential (d) Left Side Right Side Water moves to the left
Pure Water and Negative Tension Right Side Negative Water Potential 0 pressure - solute (has solutes) Water moves to the left
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Transport of Xylem Sap Pushing Xylem Guttation Root Pressure
caused by active pumping of minerals into xylem _____________: accumulation of water Guttation
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Transport of Xylem Sap ______________ Pulling Xylem Transpiration
evaporative loss of H2O from a plant through the stomata
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Transport of Xylem Sap
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The Control of Transpiration
______________ turgid - open flaccid - closed Potassium Ions active transport of H+ out of cell causes K+ to move in Guard Cells
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Stomata Open during the day / Closed at night
first light (blue light receptor) depletion of Carbon Dioxide internal clock (circadian rhythms)
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Reducing Transpiration
Small, thick leaves Thick cuticle Stomata are recessed Lose their leaves C4 or CAM plants
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Translocation of Phloem Sap
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Translocation of Phloem
Phloem Sap 30% sucrose, minerals, amino acids, hormones Transported in sieve-tube members ____________– leaves, tuber or bulbs Sugar sink – growing roots, shoots, fruits Sugar source
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Pressure Flow and Translocation
Pressure is high Pressure is low Xylem recycles water Allows Phloem sap to flow from source to sink C B
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Plant Nutrition
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Uptake of Nutrients Hydroponic
_______________ cultures used to determine which chemical elements are essential. 17 essential elements needed by all plants
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Soil Develops from weathered rocks Anchors plants Provides water
Provides dissolved minerals
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Soil Texture Pertains to sizes of soil particles
includes the following: sands ( mm) silt ( mm) clay (less than mm)
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Soil Composition Made up of sand, silt, clay, rocks, humus, microorganisms (bacteria, fungi, algae, protists, insects, worms, roots) Soil contains a mixture of different sized particles _______ – roughly equal amounts of sand, silt, and clay – most fertile Loams
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The availability of soil water and minerals
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The availability of soil water and minerals
Plant takes up water not tied to hydrophilic soil particles Positively charged ions attach to soil H+ help displace minerals attached to soil Roots add H+ to the soil directly and through the release of ____ (reacts with water to form carbonic acid) CO2
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The availability of soil water and minerals
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Soil Conservation Fertilizers (Nitrogen, Phosphorus, Potassium)
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The Nation that Destroys Its Soil Destroys Itself – Franklin D
The Nation that Destroys Its Soil Destroys Itself – Franklin D. Roosevelt 1937
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Loss of Topsoil 1930’s “_____________”
Black Blizzards 1930’s “_____________” Due to inappropriate farming in late 1800’s and early 1900’s Wheat and cattle farming Droughts Steinbeck’s Grapes of Wrath 30% of world’s farmlands have reduced production due to poor soil conditions.
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Nitrogen Fixation
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Nitrogen Fixation Plants absorb nitrogen in the form of nitrate
_____________ and ammonifying bacteria produce ammonium Ammonium is shifted to nitrate by nitrifying bacteria Plants shift nitrate back to ammonium for use Nitrogen-fixing
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Nitrogen Fixation
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Unusual Nutritional Adaptations in Plants - Epiphytes
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Unusual Nutritional Adaptations in Plants - Mistletoe
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Unusual Nutritional Adaptations in Plants – Venus Fly Traps
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Unusual Nutritional Adaptations in Plants – Pitcher Plants
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Unusual Nutritional Adaptations in Plants - Sundews
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Control Systems in Plants
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Plant Hormones Coordinates growth Coordinates development
Coordinates responses to environmental stimuli
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Plant Hormones Auxin (IAA) Cytokinins Gibberllins Abscisic Acid
Ethylene Oligogaccharins Brassinosteroids
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Auxins Stimulates stem elongation Stimulates root growth
Stimulates differentiation and branching Stimulates development of fruit Stimulates apical dominance Stimulates phototropism and gravitropism
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Auxin Control Auxin stimulates growth
Auxin block on right causes cells to elongate and the plant bends left Auxin block on left causes cells to elongate the the plant bends right
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___________ Hypothesis
Acid Growth Proton pump stimulated by auxin lower pH of wall H+ activates Enzyme Enzyme breaks hydrogen bonds in cellulose Wall takes up water and elongates
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Auxin Others Promotes secondary growth by stimulating vascular cambium and secondary xylem Promotes adventitious root at the base of a cut stem Promotes fruit growth without pollination (seedless tomatoes)
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Cytokinins Stimulates root growth
Stimulates cell division and differentiation (with auxins) more cytokinin - shoot buds develop more auxin - roots develop Stimulates germination Delays Senescence
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Gibberellins Promotes seed and bud germination
Promotes stem elongation Promotes leaf growth Stimulates flowering and fruits (with auxin)
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Abscisic Acid Slows growth Closes stomata under water stress
Permits seed dormancy
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Ethylene Promotes fruit ripening
Controls Abscission (causes leaf loss)
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Plant Movements Phototropism Gravitropism ____________ Thigmotrophism
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Plant Movement Rapid Leaf Movement (39.27)
drop in turgor pressure within pulvini sent by action potentials
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Plant Movement Sleep Movements (39.21)
cells on opposite sides of pulvinus control the movement
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Daily and Seasonal Responses
Circadian Rhythm Photoperiodism controls flowering (short-day vs. long-day) critical night length
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Photoperiodic Control
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Flowering Hormones Experiment indicates the presence of some type of flowering hormone
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Phytochromes Function as photoreceptors / red (660nm) to far red (730nm) Activates kinases (regulatory proteins)
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Red vs. Far Red Response
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Plant Responses to Environmental Stress
Water Deficit Oxygen Deprivation Salt Stress Heat Stress Cold Stress Herbivores
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Responses to Herbivores
Produce _________ (an amino acid similar to arginine) Recruitment of predatory animals Canavanine
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Why plants are important?
Food! Humans have domesticated plants for 13,000 years. ____ of all the calories consumed by humans come from six crops: Wheat, Rice, Maize, Potatoes, Cassava, and Sweet Potatoes. Also, we use plants to feed cattle, 5-7kg to produce 1 kg of beef. 80%
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Pyramid of Net Productivity
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Plants remove CO2 64
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Only 5,000 of 290,000 species have been studied.
_____ of all US Prescription Drugs contain one or more active ingredients from plants. ____ earth’s species will become extinct within the next 100 years (larger than the Permian or Cretaceous) Only 5,000 of 290,000 species have been studied. 3-4 species per hour, 27,000 per year! 25% 50%
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Cinchona tree Bark contains __________ Grows in the Andes in peru
Used since the early 1600’s to treat malaria quinine
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Aspirin Acetylsalicylic acid or ASA Dates back to 3000 B.C.
Greek Physician Hippocrates prescribed it. From _____________ and other Salicylate-rich plants (leaves and bark) Scientists at Bayer began investigating acetylsalicylic acid as a less-irritating replacement for standard common salicylate medicines. By 1899, Bayer named it this Aspirin Willow trees
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