1. Chapter 44 Table of Contents Section 1 Development of Behavior
Animal Behavior
Chapter 44
Table of Contents
Section 1 Development of Behavior
Section 2 Types of Animal Behavior

2. Section 1 Development of Behavior
Chapter 44
Objectives
Identify four questions asked by biologists who study behavior.
Describe an example of an innate behavior.
Compare four types of learned behavior.
Explain how learning and genes can interact to affect behavior.

3. Chapter 44 The Study of Behavior
Section 1 Development of Behavior
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The Study of Behavior
Ethologists are biologists who specialize in the scientific study of animal behavior.
Behavior can be defined as any action that an individual carries out in response to a stimulus or to the environment.
Two examples of behavior are a snake playing dead and a chimpanzee gathering termites on a stick.

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In order to better study behavior ethologists ask four main questions:
What causes the behavior?
What is the role of genes in the behavior?
What is the behavior’s evolutionary history?
How does the behavior affect the organism’s survival and reproduction?

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Genes and Behavior
One important factor of behavior is how certain genes affect it.
Ethologists study this by mating an animal that shows the behavior to a mate that does not.
By studying the offspring of this mating pair ethologists can see how the gene is passed on.

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Natural Selection and Behavior
Ethologists have hypothesized that animals usually behave in ways that promote their survival and offspring production.
Because genes control some behaviors, natural selection can affect genetic variation that involves behavioral genes.

7. Chapter 44 Innate Behavior
Section 1 Development of Behavior
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Innate Behavior
Some behaviors are inherited actions that are performed effectively the first time without being taught. These types of behaviors are called innate behaviors.
Fixed Action Pattern
Fixed action pattern is a type of innate behavior that all members of a species perform the same way each time they perform it.

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Innate Behavior, continued
Fixed Action Pattern, continued
Fixed action patterns continue from start to finish without modification once an environmental stimulus triggers them.
Other factors besides environmental stimulus may influence whether or not a fixed action pattern is stimulated.

9. Innate Behaviors and Roles for Bees
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Innate Behaviors and Roles for Bees
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Visual Concept

10. Chapter 44 Learned Behavior
Section 1 Development of Behavior
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Learned Behavior
When behaviors are modified by experience, it is called learning.
Four types of learning are habituation, operant conditioning, classical conditioning, and problem solving learning.
The study of learned behavior is central to much of ethology.

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Habituation
The simplest type of learning is called habituation.
Habituation is a type of learning in which an animal learns to ignore a frequent harmless stimulus.
Habituation can save energy, yet still allow for other everyday activities.

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Operant Conditioning
A type of trial and error learning is called operant conditioning, in which specific animal behaviors are deterred or reinforced by external actions upon the animal.
Operant conditioning usually happens in a controlled setting. Operant conditioning also is easier to learn if related to natural skills the animal will use for survival or reproduction.

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Classical Conditioning
Classical conditioning is a type of learning in which an animal learns to produce a specific response to a predictive stimulus in anticipation of receiving external reinforcement.
Classical conditioning can occur naturally and artificially.

14. Chapter 44 Conditioning Section 1 Development of Behavior
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Visual Concept

15. Learned Behavior, continued
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Learned Behavior, continued
Problem-Solving and Reasoning
In problem-solving learning, an animal uses several learning mechanisms, such as watching an older offspring or trial and error, to learn a type of behavior.

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Problem-Solving and Reasoning, continued
One type of problem-solving, reasoning, involves the ability to solve a problem not previously encountered by the individual in a way that is not dictated by instinct.
This type of behavior occurs without trial-and-error, as if the animal developed an insight into how to solve the problem.

17. Genes, Learning, and Behavior
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Genes, Learning, and Behavior
Most ethologists today have come to agree that animal behavior, especially complex behavior, is affected both by genes and by experience.
Imprinting
One class of behavior that is determined by both genes and learning is called imprinting. Imprinting is a form of learning in which a young animal forms permanent associations with its environment.

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Imprinting, continued
Imprinting occurs during a specific phase in an animal’s development. This period of development is called a sensitive period.
Certain forms of learning that occur during this time are very difficult to change later.

19. Section 2 Types of Animal Behavior
Chapter 44
Objectives
Discuss the optimality hypothesis and feeding behavior.
List three types of competitive behavior.
Describe three different types of reproductive behavior.
Name five kinds of communication.
Identify costs and benefits of social behavior.
Describe four types of cyclic behavior.

20. Chapter 44 Feeding Behavior
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Feeding Behavior
Animals must balance the need to obtain energy with the amount that they spend to get energy.
The optimality hypothesis is the idea that animals tend to behave in a way that maximizes food gathering, while minimizing effort and exposure to predators.

21. Chapter 44 Competitive Behavior
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Competitive Behavior
Because most resources are limited, competition for these resources occurs quite often.
There are many types of behavior that can be seen as a result of competition: aggressive behavior, territorial behavior, and dominance hierarchies.

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Aggressive Behavior
Aggressive behavior can be defined as physical conflict or threatening behavior between animals.
Aggressive behavior can be seen among males looking for a mate.

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Territorial Behavior
A territory is an area that an animal or group of animals occupies and defends from other members of the same species.
An animal will establish its territory in any number of ways and defend this area at all costs, because this helps guarantee the survival of that animal’s offspring.

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Dominance Hierarchies
Competition can lead to a clear ranking of individuals within the group, from most dominant to most subordinate. This type of ranking is called a dominance hierarchy.
This type of hierarchy reduces the need for competition and aggressive behavior as subordinates learn to submit to avoid conflict.

25. Reproductive Behavior
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Reproductive Behavior
Elaborate behaviors have evolved around the process of reproduction in many animals, which may allow these animals to recognize members of the same species or members of the opposite sex, or may even be an indicator of good health.

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Sexual Selection
Animals generally choose mates based on certain traits or behaviors, and this type of tendency is referred to as sexual selection.
These traits or behaviors will appear with increased frequency in a population, because these individuals are most likely to produce offspring.

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Sexual Selection, continued
Another means of attracting a mate involves certain behaviors and is called courtship.
In some species, courtship can include a complex series of behaviors called rituals.
A ritual is usually instinctive and may consist of specific signals and responses that indicate a willingness to mate.

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Mating Systems
Mating systems increase the likelihood that young will survive.
Male polygamy (more than one female), monogamy, and female polygamy (more than one male) are reproductive strategies that are determined primarily by the amount and type of parental care required by the young.

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Parental Behavior
The benefit of parental care is that it increases the likelihood that young will survive to adulthood.
The costs are that parental care can generally only be provided for a small number of young because of the large energy investment by the parent.

30. Nest Building Behavior
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Nest Building Behavior

31. Chapter 44 Communication
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Communication
Communication is a transfer of a signal or message from one animal to another that results in some type of response.
There are many ways animals can communicate, including sight, sound, chemicals, touch, and possibly even language.

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Sight and Sound
Color can be used to communicate certain ideas to another animals.
Bright colors often serve as a warning that an animal is poisonous. This is called aposematic coloration.
Some animals gain protection by looking like a dangerous animal. This strategy is called mimicry.
Animals can also use sound to communicate between species.

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Chemicals
Chemical communication can convey information over greater distance and time than can communication by sight or sound.
Some animals release chemicals called pheromones that cause individuals of the same species to react in a predictable way.

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Communication, continued
Touch
Species that inhabit dark hives or dens often communicate by touch in addition to using sound or chemicals.

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Language
In order to be considered language, there are certain criteria that must be met, and most animal systems are missing at least one of these criteria.
Among these are phonemes (sounds that can be combined to form words), productivity (many combinations of phonemes to produce different meanings), and grammar (rules for combining words that affect the meaning).

36. Chapter 44 Social Behavior
Section 2 Types of Animal Behavior
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Social Behavior
Social behavior can be defined as any kind of interaction between two or more animals, usually of the same species.
Some species spend the majority of their lives in social groups, others do not.

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Social Groups
Social groups have evolved in the animal kingdom because there are benefits to living in a group. These benefits can include protection from predators and more success in foraging.
There are also disadvantages to living in a social group, such as competition during courtship, theft of eggs by nonbreeding males, and possibly transmission of disease.

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Altruism
When one member of a social group acts in a way that benefits other members of the group while putting the individual at a disadvantage, this type of behavior is called altruism.
There are several ways in which an individual can be altruistic towards his social group.

39. Chapter 44 Cyclic Behavior
Section 2 Types of Animal Behavior
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Cyclic Behavior
An animal displays cyclic behaviors when it has synchronized its behavior with changes in its environment.
Biological Rhythms
There are many types of biological rhythms.
A daily biological cycle is called a circadian rhythm.

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Biological Rhythms, continued
Another type of biological rhythm is one based on the tides, and is called a lunar cycle.
There are also annual biological cycles. One such annual cycle is called hibernation.
Hibernation is a period of inactivity and lowered body temperature that some animals undergo in the winter as protection against the cold weather and lack of food.

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Migratory Behavior
Migration is a periodic group movement that is characteristic of a population or species.
Migration is exhausting and risky yet it allows animals to find habitats with plentiful seasonal foods and provides nesting sites safe from predators.

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Animal Behavior