1. Producer-Scrounger Game Role of Experience, Learning What is learned? Environmental quality (patch size) Payoffs different roles Morand-Ferron & Giraldeu (2010, Behav Ecology)

2. Producer-Scrounger Game Producer: Generates opportunities to exploit resource Finder’s share; Cost of production Scrounger : Exploits producer-generated resources Smaller share; No cost of production Exploit multiple producers

3. 0 2468101214 Probability of Energetic Failure Number of Scroungers (S) Nash Equilibrium

4. 0 2468101214 Probability of Energetic Failure Number of Scroungers (S) Nash Equilibrium

5. Producer-Scrounger Game Producer invades scrounger AND Scrounger invades producer ESS: 0 < q* < 1 where W S (q*) = W P (q*) Equilibrium predicted by game: How close? Role of learning?

6. Morand-Ferron & Giraldeau (2010) Learning Increase rate of ”correct” response Reduce errors Reduce time delay for correct response Learning the ESS: “behaviorally stable solution” Permits individual to respond to change Resource or social environment

7. Morand-Ferron & Giraldeau (2010) Learning Nutmeg mannikin P-S Interaction n-Person Game

8. Morand-Ferron & Giraldeau (2010) Aviary Flock composition fixed, 5 birds/flock Grid, 100 wells, millet seeds hidden Initial experience Low scrounging: 5 seeds in 40 wells High scrounging: 10 seeds in 20 wells Food Spatially homogeneous, heterogeneous

9. Morand-Ferron & Giraldeau (2010) Experiment: Test in minimal or high-scrounging environment 2 seeds in each well 20 seeds in 10 wells Individuals: Proportion of patches scrounged, among patches visited Nash equilibrium differed between environments Estimated from mean feeding rates High-scrounging: q* = 0.39

10. Morand-Ferron & Giraldeau (2010)

13. Discussion First demonstration learning in P-S game 1. Adjustment to environment 2. Persistent effect of experience Weigh past and recent experience Learning context can affect what is learned