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Higher Chordates: Fishes & Amphibians
Chapter 34
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Phylum Chordata
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Recall: Subphylum Urochordata Subphylum Cephalochordata Class Myxini
Tunicates Subphylum Cephalochordata Lancelets Class Myxini Hagfish Least derived craniate lineage still surviving Chordates with a head
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Subphyla Vertebrata Endoskeleton Closed circulatory system
Vertebrae: bone or cartilage Closed circulatory system Single well developed heart Skull with a brain Anterior to the spinal cord Extinct vertebrate: Conodont
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Class Agnatha Jawless fish Lack paired fins, scales
Ostracoderms extinct Cyclostomes Lampreys, hagfish Lack paired fins, scales First organisms with a backbone Appeared 500 mya
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Class Acanthodia Spiny fish Developed jaws with bony edges
Gnathostomes Internal skeletons of cartilage Extinct Appeared 430 mya
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Class Placodermi Fish with massive heads Extinct Appeared 410 mya
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Class Chondricthyes Cartilage fish (bone in teeth & scales)
Sharks, skates, rays, ratfish & chimeras Appeared 400 mya
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Chondricthyes characteristics
Premier Predators Streamlined body (power, not grace) Oil bladder (buoyancy) Acute vision (but colorblind) Detect electric fields
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Chondricthyes characteristics
Reproduction (variety) Oviparous lay eggs externally Ovoviviparous keep eggs in uterus until hatching Viviparous Placenta – live birth
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Class Osteichthyes Ossification of the skeleton (Bone)
Draw H2O across gills Operculum bony covering Swim bladder (buoyancy) Mostly oviparous Appeared 400 mya 95% of today’s fishes
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Subclass Actinopterygii
Ray finned fish Bony rays in fins for maneuvering, defense, and other functions
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Subclass Sarcopterygii
Lobe finned fish Rod-shaped bones in fins surrounded with thickened layer of muscle Swim or “walk” across substrate Actinista (coelacanth) Dipnoi (lungfish)
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Evolution of tetrapods
One of the most significant events in vertebrate history was when the fins of some lobe-finned organism evolved into the limbs and feet of tetrapods
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Class Amphibia “Two lives” Gas exchange
aquatic larva develops into a terrestrial adult Gas exchange Lungs and moist skin All amphibians must lay eggs in water
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Order Urodela Salamanders tails
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Order Anura Frogs and toads No tail
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Order Apoda Caecilians Legless Wormlike Burrowing
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Amphibian Metamorphosis
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