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Ecosystems and Energy Structure
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Ecosystems An ecosystem comprises a community and its physical environment Both biotic and abiotic components make up an ecosystem
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Energy Flow and Chemical Cycling
One way to look at ecosystems is as systems that cycle life-sustaining chemicals and allow energy to flow between trophic levels Trophic levels describe the levels of the food chain
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Energy Flow All energy used for life must originally come from autotrophs Autotrophs obtain carbon from inorganic compounds Photoautotrophs obtain energy from light Chemoautotrophs obtain energy from inorganic compounds
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Primary production Autotrophs are responsible for primary production, the creation of sugars and other energy-rich compounds from inorganic matter
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Heterotrophs Heterotrophs obtain carbon from organic sources
Chemoheterotrophs obtain energy from sources such as carbohydrates Photoheterotrophs obtain energy from light
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Consumers Heterotrophs act as consumers
Organisms that eat primary producers are primary consumers Organisms that eat primary consumers are secondary consumers, and so on (tertiary, quaternary)
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Decomposition Decomposers (also called saprotrophs) include fungi, bacteria. Primary difference between decomposers and consumers is that decomposers break down dead matter while consumers eat still living or recently deceased matter. Similar to detritivore
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Decomposers play a vital part in chemical recycling, allowing producers to continue producing.
Fungi (rot) and some bacteria are the only known organisms able to digest lignin, a primary component of wood.
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Laws of Physics The laws of physics still apply when dealing with biology Of particular importance are the Laws of Thermodynamics and Conservation of Mass
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Laws of Thermodynamics
1st – Energy cannot be created or destroyed, only transformed. 2nd – Energy transformations are never completely efficient. Combined the two laws have a variety of implications for life. Life could not exist without the constant addition of energy, primarily from the Sun, to our ecosystems.
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Law of Conservation of Mass
Mass cannot be created or destroyed, only converted into new forms For ecosystems the implication of this law is that carbon, nitrogen, phosphorus, and other nutrients cannot be destroyed and will be continuously recycled
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Primary Production The sun provides approximately 1022 J (2.39 x 1021 cal) of energy to the Earth per day However most of this energy is absorbed or deflected by Earth’s atmosphere, or land places without any photosynthetic organism around In the end, approximately 1% actually makes it into chemical energy
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Gross vs Net Gross primary productivity (GPP) is the total about of light energy converted into chemical energy in a given time period Net Primary Productivity (NPP) = GPP – R (energy used by primary producers for respiration) NPP is the amount of energy that is available to consumers
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Measuring energy production
Most commonly energy production is measured by biomass added to the ecosystem in a given time Note that it’s the biomass added. The total biomass (called the standing crop) is not a useful indicator of energy production,
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What limits productivity?
The limiting nutrient is the nutrient that most limits primary production Put another way, the limiting nutrient is the nutrient that most increases production when added to an ecosystem
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The limiting nutrient depends upon the ecosytem in question
In ocean ecosystems the limiting nutrients tend to be either light or nitrogen Light limits productivity in deep water where little light penetrates Near the ocean surface nutrients, such as nitrogen, limit productivity
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Nitrogen is the most common limiting nutrient in open water
Addition of nitrogen to open water often results in eutrophication and algal blooms
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Upwelling can cause natural increase in nutrients
Wind driven motion of water can cause nutrient rich upwellings
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Freshwater tends to lack phosphorus
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Terrestrial ecosystems
Water can be a very important limiting nutrient in terrestrial systems Temperature also varies more on land than in water and can limit growth in colder months Nitrogen and phosphorus may be local limiters on productivity.
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