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Photosynthesis: Using Light to Make Food

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1 Photosynthesis: Using Light to Make Food
Chapter 7 Photosynthesis: Using Light to Make Food

2 Biology and Society: Biofuels
Wood has historically been the main fuel used to produce heat and light. © 2013 Pearson Education, Inc. 2

3 Figure 7.0 Figure 7.0 Capturing solar energy 3

4 Biology and Society: Biofuels
Industrialized societies replaced wood with fossil fuels including coal, gas, and oil. To limit the damaging effects of fossil fuels, researchers are investigating the use of biomass (living material) as efficient and renewable energy sources. © 2013 Pearson Education, Inc. 4

5 Biology and Society: Biofuels
There are several types of biofuels. Bioethanol is a type of alcohol produced by the fermentation of glucose made from starches in crops such as grains, sugar beets, and sugar cane. Bioethanol may be used directly as a fuel source in specially designed vehicles or as a gasoline additive. © 2013 Pearson Education, Inc. 5

6 Biology and Society: Biofuels
Cellulosic ethanol is a type of bioethanol made from cellulose in nonedible plant material such as wood or grass. Biodiesel is made from plant oils or recycled frying oil. © 2013 Pearson Education, Inc. 6

7 THE BASICS OF PHOTOSYNTHESIS
is used by plants, algae (protists), and some bacteria, transforms light energy into chemical energy, and uses carbon dioxide and water as starting materials. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters. 7

8 THE BASICS OF PHOTOSYNTHESIS
The chemical energy produced via photosynthesis is stored in the bonds of sugar molecules. Organisms that use photosynthesis are photosynthetic autotrophs and the producers for most ecosystems. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters.

9 Photosynthetic Protists Photosynthetic Bacteria
Figure 7.1 PHOTOSYNTHETIC AUTOTROPHS Plants (mostly on land) Photosynthetic Protists (aquatic) Photosynthetic Bacteria (aquatic) LM Forest plants Kelp, a large, multicellular alga Micrograph of cyanobacteria Figure 7.1 A diversity of photosynthetic autotrophs 9

10 Plants (mostly on land)
Figure 7.1a Plants (mostly on land) Forest plants Figure 7.1 A diversity of photosynthetic autotrophs (part 1) 10

11 Photosynthetic Protists
Figure 7.1b Photosynthetic Protists (aquatic) Kelp, a large, multicellular alga Figure 7.1 A diversity of photosynthetic autotrophs (part 2) 11

12 Photosynthetic Bacteria
Figure 7.1c Photosynthetic Bacteria (aquatic) LM Micrograph of cyanobacteria Figure 7.1 A diversity of photosynthetic autotrophs (part 3) 12

13 Chloroplasts: Sites of Photosynthesis
Chloroplasts are the site of photosynthesis and found mostly in the interior cells of leaves. Inside chloroplasts are interconnected, membranous sacs called thylakoids, which are suspended in a thick fluid called stroma. Thylakoids are concentrated in stacks called grana. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters. 13

14 Chloroplasts: Sites of Photosynthesis
The green color of chloroplasts is from chlorophyll, a light-absorbing pigment that plays a central role in converting solar energy to chemical energy. Stomata are tiny pores in leaves where carbon dioxide enters and oxygen exits. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters. 14

15 15 Photosynthetic cells Vein Stomata Leaf cross section Figure 7.2-1
CO2 O2 Stomata Leaf cross section Figure 7.2 Journey into a leaf (step 1) 15

16 16 Chloroplast Photosynthetic cells Vein Stomata Leaf cross section
Figure 7.2-2 Photosynthetic cells Chloroplast Vein CO2 O2 Stomata LM Leaf cross section Interior cell Figure 7.2 Journey into a leaf (step 2) 16

17 17 Chloroplast Photosynthetic cells Inner and outer membranes Vein
Figure 7.2-3 Photosynthetic cells Chloroplast Inner and outer membranes Vein Thylakoid space Stroma Granum CO2 O2 Stomata LM Leaf cross section Interior cell Colorized TEM Figure 7.2 Journey into a leaf (step 3) 17

18 18 Vein (transports water and nutrients) Photosynthetic cells Stomata
Figure 7.2a Vein (transports water and nutrients) Photosynthetic cells CO2 O2 Stomata Leaf cross section Figure 7.2 Journey into a leaf (part 1) 18

19 19 Chloroplast Inner and outer membranes Thylakoid Stroma space Granum
Figure 7.2b Chloroplast Inner and outer membranes Thylakoid space Stroma Granum LM Interior cell Colorized TEM Figure 7.2 Journey into a leaf (part 2) 19

20 20 Chloroplast Interior cell LM Figure 7.2c
Figure 7.2 Journey into a leaf (part 3) 20

21 21 Stroma Granum Colorized TEM Figure 7.2d
Figure 7.2 Journey into a leaf (part 4) 21

22 The Simplified Equation for Photosynthesis
In the overall equation for photosynthesis, notice that the reactants of photosynthesis are the waste products of cellular respiration. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters. 22

23 23 Light energy 6 CO2 6 H2O C6H12O6 6 O2 Photo- synthesis Carbon
Figure 7-UN01 Light energy 6 CO2 6 H2O C6H12O6 6 O2 Photo- synthesis Carbon dioxide Water Glucose Oxygen gas In-text figure, leaf with photosynthesis equation, p. 109 23

24 The Simplified Equation for Photosynthesis
In photosynthesis, sunlight provides the energy, electrons are boosted “uphill” and added to carbon dioxide, and sugar is produced. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters. 24

25 The Simplified Equation for Photosynthesis
During photosynthesis, water is split into hydrogen and oxygen. Hydrogen is transferred along with electrons and added to carbon dioxide to produce sugar. Oxygen escapes through stomata into the atmosphere. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters. 25

26 A Photosynthesis Road Map
Photosynthesis occurs in two multistep stages: the light reactions convert solar energy to chemical energy and the Calvin cycle uses the products of the light reactions to make sugar from carbon dioxide. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters. 26

27 A Photosynthesis Road Map
The initial incorporation of carbon from the atmosphere into organic compounds is called carbon fixation. This lowers the amount of carbon in the air. Deforestation reduces the ability of the biosphere to absorb carbon by reducing the amount of photosynthetic plant life. BioFlix Animation: Photosynthesis © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Students often do not fully understand how the burning of fossil fuels contributes to global warming. They might wonder, “How does the burning of fossil fuels differ from the burning of ethanol produced from crops?” Students might not realize that the carbon in fossil fuels was removed from the atmosphere hundreds of millions of years ago, while the carbon in crops was removed much more recently, when the crops were grown. The use of ethanol as an alternative is complicated by the typical reliance upon fossil fuels for ethanol production. 2. Some students do not realize that plant cells also have mitochondria. Instead, they assume that the chloroplasts are sufficient for the plant cell’s needs. But nearly 50% of the carbohydrates produced by plant cells are used for cellular respiration (involving mitochondria). 3. Students may not connect the growth in plant mass to the fixation of carbon during the Calvin cycle. It can be difficult for many students to appreciate that molecules in air can contribute significantly to the mass of plants. 4. Students may understand the overall chemical relationships between photosynthesis and cellular respiration, but many struggle to understand the use of carbon dioxide in the Calvin cycle. Photosynthesis is much more than gas exchange. 5. Students who have not read all of Chapter 7 may not realize that glucose is not the direct product of photosynthesis. Although glucose is shown as a product of photosynthesis, a three-carbon sugar is directly produced (G3P). A plant can use G3P to make many types of organic molecules, including glucose. (The authors address the production of G3P under the section “The Calvin Cycle” later in this chapter.) Teaching Tips 1. When introducing the diverse ways that plants impact our lives, consider challenging your students to come up with a list of products made from plants that they encounter regularly. Perhaps you might only list those encountered in a single day of college life. The list can be surprising and help to build up your “catalog of examples.” 2. The living world contains many examples of adaptations to increase surface area. Some examples are the many folds of the inner mitochondrial membrane, the highly branched surfaces of fish gills and human lungs, and the highly branched system of capillaries in the tissues of our bodies. Consider relating this broad principle seen elsewhere to the extensive folding of the thylakoid membranes. 3. In our world, energy is frequently converted to a usable form in one place and used in another. For example, electricity is generated by power plants, transferred to our homes, and used to run computers, create light, and help us prepare foods. Consider relating this common energy transfer to the two-stage process of photosynthesis. 4. You might wish to discuss the evolution of chloroplasts from photosynthetic prokaryotes if you will not address this subject elsewhere in your course. 5. Figure 7.3 is an important visual organizer that notes the key structures and functions of the two stages of photosynthesis. This figure reminds students where water and sunlight are used in the thylakoid membranes to generate oxygen, ATP, and NADPH. The second step, in the stroma, reveals the use of carbon dioxide, ATP, and NADPH to generate carbohydrates. 6. The thylakoid space and the intermembrane space of a mitochondrion have analogous roles. Students might be encouraged to create a list of the similarities in structure and function of mitochondria and chloroplasts through these related chapters. 27

28 28 H2O Chloroplast Light Light reactions ATP – – NADPH O2 Figure 7.3-1
Figure 7.3 A road map for photosynthesis (step 1) 28

29 29 H2O CO2 Chloroplast Light NADP+ ADP  P Calvin cycle Light
Figure 7.3-2 H2O CO2 Chloroplast Light NADP+ ADP P Calvin cycle Light reactions ATP NADPH O2 Sugar Figure 7.3 A road map for photosynthesis (step 2) 29

30 THE LIGHT REACTIONS: CONVERTING SOLAR ENERGY TO CHEMICAL ENERGY
Chloroplasts are chemical factories powered by the sun and convert sunlight into chemical energy. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 30

31 Calvin cycle Light reactions
Figure 7-UN02 CO2 H2O Light NADP ADP + P Calvin cycle Light reactions ATP NADPH O2 Sugar In-text figure, light reactions, p. 110 31

32 The Nature of Sunlight Sunlight is a type of energy called radiation, or electromagnetic energy. The distance between the crests of two adjacent waves is called a wavelength. The full range of radiation is called the electromagnetic spectrum. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 32

33 33 10–5 nm 10–3 nm 1 nm 103 nm 106 nm 1 m 103 m Increasing wavelength
Figure 7.4 Increasing wavelength 10–5 nm 10–3 nm 1 nm 103 nm 106 nm 1 m 103 m Gamma rays Micro- waves Radio waves X-rays UV Infrared Visible light 380 400 500 600 700 750 Wavelength (nm) Wavelength  580 nm Figure 7.4 The electromagnetic spectrum 33

34 34 Light Reflected light Chloroplast Absorbed light Transmitted
Figure 7.5 Light Reflected light Chloroplast Absorbed light Transmitted light (detected by your eye) Figure 7.5 Why are leaves green? 34

35 35 Light Reflected light Chloroplast Absorbed light Transmitted
Figure 7.5a Light Reflected light Chloroplast Absorbed light Transmitted light (detected by your eye) Figure 7.5 Why are leaves green? (part 1) 35

36 Figure 7.5b Figure 7.5 Why are leaves green? (part 2) 36

37 The Process of Science: What Colors of Light Drive Photosynthesis?
Observation: In 1883, German biologist Theodor Engelmann saw that certain bacteria tend to cluster in areas with higher oxygen concentrations. Question: Could this information determine which wavelengths of light work best for photosynthesis? Hypothesis: Oxygen-seeking bacteria will congregate near regions of algae performing the most photosynthesis. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 37

38 The Process of Science: What Colors of Light Drive Photosynthesis?
Experiment: Engelmann laid a string of freshwater algal cells in a drop of water on a microscope slide, added oxygen-sensitive bacteria to the drop, and used a prism to create a spectrum of light shining on the slide. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 38

39 Animation: Light and Pigments
The Process of Science: What Colors of Light Drive Photosynthesis? Results: Bacteria mostly congregated around algae exposed to red-orange and blue-violet light and rarely moved to areas of green light. Conclusion: Chloroplasts absorb light mainly in the blue-violet and red-orange part of the spectrum. Animation: Light and Pigments © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 39

40 40 Light Prism Microscope slide Bacteria Bacteria Number of bacteria
Figure 7.6 Light Prism Microscope slide Bacteria Bacteria Number of bacteria Algal cells 400 500 600 700 Wavelength of light (nm) Figure 7.6 Investigating how light wavelength affects photosynthesis 40

41 Chloroplast Pigments Chloroplasts contain several pigments:
Chlorophyll a absorbs mainly blue-violet and red light and participates directly in the light reactions. Chlorophyll b absorbs mainly blue and orange light and participates indirectly in the light reactions. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 41

42 Chloroplast Pigments Carotenoids
absorb mainly blue-green light, participate indirectly in the light reactions, and absorb and dissipate excessive light energy that might damage chlorophyll. The spectacular colors of fall foliage are due partly to the yellow-orange light reflected from carotenoids. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 42

43 Figure 7.7 Figure 7.7 Photosynthetic pigments 43

44 How Photosystems Harvest Light Energy
Light behaves as photons, a fixed quantity of light energy. Chlorophyll molecules absorb photons. Electrons in the pigment gain energy. As the electrons fall back to their ground state, energy is released as heat or light. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 44

45 45 (a) Absorption of a photon (b) Fluorescence of a glow stick
Figure 7.8 Excited state Absorption of a photon excites an electron. The electron falls to its ground state. e– Heat Light Light (fluorescence) Photon Ground state Chlorophyll molecule (a) Absorption of a photon (b) Fluorescence of a glow stick Figure 7.8 Excited electrons in pigments 45

46 (a) Absorption of a photon
Figure 7.8a Excited state Absorption of a photon excites an electron. The electron falls to its ground state. e– Heat Light Light (fluorescence) Photon Ground state Chlorophyll molecule (a) Absorption of a photon Figure 7.8 Excited electrons in pigments (part 1) 46

47 (b) Fluorescence of a glow stick
Figure 7.8b (b) Fluorescence of a glow stick Figure 7.8 Excited electrons in pigments (part 2) 47

48 How Photosystems Harvest Light Energy
In the thylakoid membrane, chlorophyll molecules are organized with other molecules into photosystems. A photosystem is a cluster of a few hundred pigment molecules that function as a light-gathering antenna. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 48

49 How Photosystems Harvest Light Energy
The reaction center of the photosystem consists of chlorophyll a molecules that sit next to another molecule called a primary electron acceptor, which traps the light-excited electron from chlorophyll a. Another team of molecules built into the thylakoid membrane then uses that trapped energy to make ATP and NADPH. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 49

50 50 Figure 7.9 Chloroplast Cluster of pigment molecules Photon Primary
electron acceptor e– Electron transfer Reaction center Reaction- center chlorophyll a Pigment molecules Thylakoid membrane Transfer of energy Photosystem Figure 7.9 A photosystem: light-gathering molecules that focus light energy onto a reaction center 50

51 51 Chloroplast Cluster of pigment molecules Thylakoid membrane
Figure 7.9a Chloroplast Cluster of pigment molecules Thylakoid membrane Figure 7.9 A photosystem: light-gathering molecules that focus light energy onto a reaction center (part 1) 51

52 52 Photon Primary electron acceptor Electron transfer Reaction center
Figure 7.9b Photon Primary electron acceptor Electron transfer e– Reaction center Reaction- center chlorophyll a Pigment molecules Transfer of energy Photosystem Figure 7.9 A photosystem: light-gathering molecules that focus light energy onto a reaction center (part 2) 52

53 How the Light Reactions Generate ATP and NADPH
Two types of photosystems cooperate in the light reactions: the water-splitting photosystem and the NADPH-producing photosystem. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 53

54 54 1 Water-splitting photosystem Primary electron acceptor Light
Figure Primary electron acceptor 2e Light 1 Reaction- center chlorophyll H2O 2e Water-splitting photosystem 2 H O2 Figure 7.10 The light reactions of photosynthesis (step 1) 54

55 55 2 1 Water-splitting photosystem Energy to make ATP Primary electron
Figure Energy to make ATP Primary electron acceptor 2 2e Electron transport chain Light 1 Reaction- center chlorophyll H2O 2e Water-splitting photosystem 2 H O2 Figure 7.10 The light reactions of photosynthesis (step 2) 55

56 56 3 2 1 NADPH-producing photosystem Water-splitting photosystem
Figure Primary electron acceptor NADP 2e Energy to make 3 ATP Primary electron acceptor 2e 2 NADPH 2e Light Electron transport chain Light Reaction- center chlorophyll 1 Reaction- center chlorophyll NADPH-producing photosystem H2O 2e Water-splitting photosystem 2 H O2 Figure 7.10 The light reactions of photosynthesis (step 3) 56

57 How the Light Reactions Generate ATP and NADPH
The light reactions are located in the thylakoid membrane. An electron transport chain connects the two photosystems and releases energy that the chloroplast uses to make ATP. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. The authors note that sunlight is a type of radiation. Many students think of radiation as a result of radioactive decay, a serious threat to health. The diverse types of radiation and the varying energy associated with each, might need to be explained. 2. The authors note that electromagnetic energy travels through space in waves that are like those made by a pebble dropped in a pond. This wave imagery is helpful, but can confuse students when energy is also thought of as discrete packets called photons. The dual nature of light as a wave and particle may need to be discussed further, if students are to do more than just accept the definitions. 3. The light reactions are not solely responsible for all of the products in the general chemical equation of photosynthesis. Point out that oxygen is generated in the light reactions but that glucose results from products of the Calvin cycle. 4. Even at the college level, students struggle to understand why we perceive certain colors. The authors discuss the specific absorption and reflection of certain wavelengths of light, noting which colors are absorbed and which are reflected (and thus available for our eyes to detect). Consider spending time to make sure that your students understand how photosynthetic pigments absorb and reflect certain wavelengths. Teaching Tips 1. Consider bringing a prism to class and demonstrating the spectrum of visible light. Depending on what you have available, it can be a dramatic reinforcement of this key concept. You might show an image of a rainbow if you are willing to explain how it is produced. 2. Consider bringing a glow stick to class just to help illustrate the authors’ note about the light-generating reactions in glow sticks. Small demonstrations break up lectures and stir attention. 3. The authors discuss a phenomenon that most students have noticed: Dark surfaces heat up faster in the sun than lighter-colored surfaces. This is an opportunity to demonstrate to your students the various depths of scientific explanations and help them appreciate their own educational progress. In elementary school, they might have learned that the sun heats darker surfaces faster than lighter surfaces. In high school, they may have learned about light energy and the fact that dark surfaces absorb more of this energy than lighter surfaces. Now, in college, they are learning that at the atomic level, darker surfaces absorb the energy of more photons, exciting more electrons, which then fall back to a lower state, releasing more heat. 4. The authors develop a mechanical analogy to the energy levels and movement of electrons in the light reaction. Figure 7.12 equates the height of an electron with its energy state. Thus, electrons captured at high levels carry more energy than electrons in lower positions. Although this figure can be very effective, students might need to be carefully led through the analogy to understand precisely what is represented. 57

58 58 – Figure 7.11 To Calvin cycle Light Light Stroma Thylakoid ATP
NADPH ATP ADP P NADP H Stroma Electron transport chain Thylakoid membrane Photosystem Photosystem ATP synthase Inside thylakoid Electron flow 2e H H H2O H H H O2 Thylakoid membrane Figure 7.11 How the thylakoid membrane converts light energy to the chemical energy of NADPH and ATP 58

59 59 Stroma Thylakoid membrane Inside thylakoid Thylakoid membrane H
Figure 7.11a H Stroma Electron transport chain Thylakoid membrane Photosystem Inside thylakoid 2e H2O H O2 Thylakoid membrane Figure 7.11 How the thylakoid membrane converts light energy to the chemical energy of NADPH and ATP (part 1) 59

60 60 To Calvin cycle Light ATP synthase Electron flow H NADPH ATP ADP 
Figure 7.11b To Calvin cycle Light H NADPH ATP ADP P NADP H Electron transport chain Photosystem ATP synthase Electron flow H H H H H Figure 7.11 How the thylakoid membrane converts light energy to the chemical energy of NADPH and ATP (part 2) 60

61 61 – – – – – – – – – ATP NADPH Water-splitting photosystem
Figure 7.12 e ATP e e NADPH e e e Photon e Photon Water-splitting photosystem NADPH-producing photosystem Figure 7.12 A hard-hat analogy for the light reactions 61

62 THE CALVIN CYCLE: MAKING SUGAR FROM CARBON DIOXIDE
functions like a sugar factory within a chloroplast and regenerates the starting material with each turn. Blast Animation: Photosynthesis: Light-Independent Reactions © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Glucose is not the direct product of the Calvin cycle, as might be expected from the general equation for photosynthesis. Instead, G3P, as noted in the text, is the main product. Clarify for students the diverse uses of G3P in the production of many important plant molecules and the advantages of producing a molecule with this flexibility. Teaching Tips 1. If you can find examples of potted C3, C4, and CAM plants, consider bringing them to class. Referring to living plants in class helps students understand these abstract concepts. Nice photographs can serve as a substitute. 2. Relate the properties of C3 and C4 plants to the regions of the country where each is grown. Students might generally understand that crops have specific requirements, but may not specifically relate these physiological differences to their geographic sites of production or specific evolutionary histories. 62

63 Calvin cycle Light reactions
Figure 7-UN03 CO2 Light H2O NADP ADP + P Calvin cycle Light reactions ATP NADPH O2 Sugar In-text figure, Calvin cycle, p. 115 63

64 64 1 CO2 (from air) RuBP sugar Three-carbon molecule Calvin cycle P P
Figure CO2 (from air) 1 P RuBP sugar Three-carbon molecule P P Calvin cycle Figure 7.13 The Calvin cycle (step 1) 64

65 65 1 2 CO2 (from air) RuBP sugar Three-carbon molecule Calvin cycle
Figure CO2 (from air) 1 P RuBP sugar Three-carbon molecule ATP P P ADP P Calvin cycle NADPH NADP G3P sugar P 2 Figure 7.13 The Calvin cycle (step 2) 65

66 Glucose (and other organic
Figure CO2 (from air) 1 P RuBP sugar Three-carbon molecule ATP P P ADP P Calvin cycle NADPH NADP G3P sugar G3P sugar 3 P P 2 G3P sugar Glucose (and other organic compounds) P Figure 7.13 The Calvin cycle (step 3) 66

67 Glucose (and other organic
Figure CO2 (from air) 1 P RuBP sugar Three-carbon molecule 4 ATP P P ADP P ADP P Calvin cycle NADPH ATP NADP G3P sugar G3P sugar 3 P P 2 G3P sugar Glucose (and other organic compounds) P Figure 7.13 The Calvin cycle (step 4) 67

68 Evolution Connection: Solar-Driven Evolution
C3 plants use CO2 directly from the air and are very common and widely distributed. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Glucose is not the direct product of the Calvin cycle, as might be expected from the general equation for photosynthesis. Instead, G3P, as noted in the text, is the main product. Clarify for students the diverse uses of G3P in the production of many important plant molecules and the advantages of producing a molecule with this flexibility. Teaching Tips 1. If you can find examples of potted C3, C4, and CAM plants, consider bringing them to class. Referring to living plants in class helps students understand these abstract concepts. Nice photographs can serve as a substitute. 2. Relate the properties of C3 and C4 plants to the regions of the country where each is grown. Students might generally understand that crops have specific requirements, but may not specifically relate these physiological differences to their geographic sites of production or specific evolutionary histories. 68

69 Evolution Connection: Solar-Driven Evolution
C4 plants close their stomata to save water during hot and dry weather and can still carry out photosynthesis. CAM plants are adapted to very dry climates and open their stomata only at night to conserve water. © 2013 Pearson Education, Inc. Student Misconceptions and Concerns 1. Glucose is not the direct product of the Calvin cycle, as might be expected from the general equation for photosynthesis. Instead, G3P, as noted in the text, is the main product. Clarify for students the diverse uses of G3P in the production of many important plant molecules and the advantages of producing a molecule with this flexibility. Teaching Tips 1. If you can find examples of potted C3, C4, and CAM plants, consider bringing them to class. Referring to living plants in class helps students understand these abstract concepts. Nice photographs can serve as a substitute. 2. Relate the properties of C3 and C4 plants to the regions of the country where each is grown. Students might generally understand that crops have specific requirements, but may not specifically relate these physiological differences to their geographic sites of production or specific evolutionary histories. 69

70 70 C4 Pathway CAM Pathway (example: sugarcane) (example: pineapple)
Figure 7.14 ALTERNATIVE PHOTOSYNTHETIC PATHWAYS C4 Pathway (example: sugarcane) CAM Pathway (example: pineapple) Cell type 1 CO2 CO2 Night Four-carbon compound Four-carbon compound Cell type 2 CO2 CO2 Calvin cycle Calvin cycle Sugar Sugar Day C4 plant CAM plant Figure 7.14 C4 and CAM photosynthesis 70

71 C4 Pathway (example: sugarcane) CAM Pathway (example: pineapple)
Figure 7.14a C4 Pathway (example: sugarcane) CAM Pathway (example: pineapple) Cell type 1 CO2 CO2 Night Four-carbon compound Four-carbon compound Cell type 2 CO2 CO2 Calvin cycle Calvin cycle Sugar Sugar Day CAM plant C4 plant Figure 7.14 C4 and CAM photosynthesis (part 1) 71

72 C4 Pathway (example: sugarcane)
Figure 7.14b C4 Pathway (example: sugarcane) Figure 7.14 C4 and CAM photosynthesis (part 2) 72

73 CAM Pathway (example: pineapple)
Figure 7.14c CAM Pathway (example: pineapple) Figure 7.14 C4 and CAM photosynthesis (part 3) 73

74 74 Light energy Photosynthesis Carbon dioxide Water Glucose Oxygen gas
Figure 7-UN04 Light energy 6 CO2 6 H2O C6H12O6 6 O2 Photosynthesis Carbon dioxide Water Glucose Oxygen gas Summary of Key Concepts: The Simplified Equation for Photosynthesis 74

75 75 Chloroplast CO2 Light H2O Stack of thylakoids NADP Stroma ADP P
Figure 7-UN05 Chloroplast CO2 Light H2O Stack of thylakoids NADP Stroma ADP P Calvin cycle Light reactions ATP NADPH Sugar used for cellular respiration cellulose starch other organic compounds O2 Sugar Summary of Key Concepts: A Photosynthesis Road Map 75

76 76 + NADP e – 2e – ADP acceptor ATP e – 2e – acceptor – – 2e – NADPH
Figure 7-UN06 NADP e 2e ADP acceptor ATP e 2e acceptor 2e NADPH Photon Electron transport chain Photon Chlorophyll H2O Chlorophyll NADPH-producing photosystem 2e Water-splitting photosystem 2 1 2 H O2 + Summary of Key Concepts: How Photosystems Harvest Light Energy; How the Light Reactions Generate ATP and NADPH 76

77 77 CO2 Calvin cycle ADP NADP Glucose and other compounds
Figure 7-UN07 CO2 ATP ADP P Calvin cycle NADPH NADP G3P Glucose and other compounds (such as cellulose and starch) P Summary of Key Concepts: The Calvin Cycle: Making Sugar from Carbon Dioxide 77


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