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Principles of evolution
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evolution The process of biological change by which descendants come to differ from their ancestors Early scientists proposed ideas about evolution
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Carolus linnaeus 1700s Swedish botanist
Developed a classification system for all organisms known at the time based on their similarities and evolutionary relationships Binomial Nomenclature
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Georges louis leclerc de buffon
1700s French Naturalist Proposed that species shared common ancestors instead of arising separately Challenged the idea that Earth was only 6,000 years old Ideas were similar to geologist Charles Lyell’s
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Erasmus darwin Darwin’s grandfather, born in 1731
Respected English doctor and poet Proposed that: all living things were descended from a common ancestor more complex forms of life arose from less-complex life forms
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Jean-baptiste lamarck
French naturalist Proposed that all organisms evolved toward perfection and complexity
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Jean Baptiste Lamarck Believed that over a lifetime of an individual physical features: Increase in size the more they are used Decrease in size the less they are used These changes are then passed onto offspring “Inheritance of Acquired Characteristics”
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Lamarck’s Theory was… WRONG!
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Charles Darwin ( ) 1859: “On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life” Has convincing evidence that species evolve Describes a reasonable explanation of how evolution occurs
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Charles Darwin Destined to become a doctor or a minister
Liked the outdoors and collecting samples Age 22: invited to be a naturalist on a voyage of the HMS Beagle History would never be the same
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Darwin’s Voyage Charles Lyell’s book: “Principles of Geology”
Book suggested the surface of the Earth changed slowly over many years Darwin thought: couldn’t species have changed too?
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Thomas Malthus English economist
1798 essay claimed human populations are able to increase faster than the food demand They do not grow unchecked because death caused by disease, war and famine slows population growth Darwin thought: Doesn’t this happen with all living species?
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Theories of geologic change set the stage for darwin’s theory
Fossil: trace of an organism from the past Catastrophism: theory that states that natural disasters such as floods and volcanic eruptions shaped Earth’s landforms and caused extinction of some species Gradualism: principle that states that the changes in landforms result from slow changes over a long period of time Uniformitarianism: theory that states that the geological processes that shape Earth are uniform through time
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Theories of geologic change set the stage for darwin’s theory
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The Fossil Record Fossils of animals show a pattern of development from early ancestors to modern descendants Offer the most direct evidence that evolution takes place
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The Fossil Record Darwin predicted the intermediate forms between the great groups of organisms that would be found Darwin’s theory is almost universally accepted by scientists as the best available explanation for biological diversity on Earth Fossil intermediaries have been found between: Fishes and amphibians Reptiles and birds Reptiles and mammals A model of Tiktaalik roseae, which fills in the evolutionary gap between fish and land animals, is depicted in what scientists believe to be the animal's environment about 375 million years ago.
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The Fossil Record Most scientists agree:
Earth is about 4.5 billion years old Organisms have inhabited Earth for most of its history All organisms living today share common ancestry with earlier, simpler life-forms The fossil record is not complete The record of evolution of life is not complete
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Darwin’s Voyage Darwin saw things in his travels that could only be explained by gradual change Visited the Galapagos Islands, 1000 km off the coast of Ecuador
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Darwin’s Voyage “Descent With Modification”-Evolution
Noticed many of the plants and animals of the Galapagos resembled those of the nearby coast of S. America Suggested the species migrated from S. America and changed after they arrived
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Darwin’s observations
Darwin observed differences among island species Variation: differences in physical traits of an individual from the group to which it belongs Adaptation: inherited trait that is selected for over time because it allows organisms to better survive in their environment
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Darwin’s Finches 9 distinct species of finches in the Galapagos Islands, all very similar to each other except for their bills 2 ground finches: large bills to feed on seeds that they crush with their beaks 2 with narrower bills eat insects 1 finch eats fruit 1 picks insects out of cactuses 1 drinks the blood of sea birds
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Darwin’s Finches Darwin suggested that the nine species evolved from a common ancestral species During a year of rain, food is plentiful for all the finches
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Darwin’s Finches Dry years Wet years
Longer, more massive beaks Wet years Smaller beaks returned The number of birds with different beak shapes are changed by natural selection in response to the available food supply, as Darwin suggested
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Darwin observed fossil and geologic evidence supporting an ancient earth
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Publication of Darwin’s Work
1844 Darwin wrote down his ideas Due to public criticism of Lamarck’s hypotheses, Darwin did not publish his findings 1858 Alfred Russel Wallace wrote an essay that summarized exactly what Darwin had believed for years! 1859 Darwin quickly published his book
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Darwin’s Theory Inherited variation exists within the genes of every population or species The result of random mutation and translation errors
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Darwin’s Theory 2. In a particular environment, some individuals of a population or species are better suited to survive A result of variation Have more offspring
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Darwin’s Theory 3. Over time, the traits that make certain individuals of a population able to survive and reproduce tend to spread in that population
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Darwin’s Theory 4. There is overwhelming evidence from fossils and many other sources that living species evolved from organisms that are extinct
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Several key insights led to Darwin's idea for natural selection
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Artificial selection Process by which humans modify a species by breeding it for certain traits
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Natural Selection Individuals that have physical or behavioral traits that better suit their environment More likely to survive More likely to reproduce successfully Peppered Moth Example
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Natural Selection The number of individuals that carry favorable characteristics that are inherited will increase in the population The nature of the population will change… Evolution
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Population All the individuals of a species that live in a specific geographical area and that can interbreed
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Natural selection explains how evolution can occur
Variation Overproduction Adaptation Descent with modification
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principles of natural selection
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fitness Measure of an organism’s ability to survive and produce offspring relative to other members of a population s
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Natural selection acts on existing variation
Acts on phenotypes, not genotypes
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Evidence of evolution in Darwin’s time came from several sources
Fossils Geography Embryology
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Homologous Structures
Anatomical structures that share a common ancestry
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Homologous Structures
Most scientists believe that the evolutionary history of organisms is also seen in the development of embryos At some point, all vertebrate embryos have a tail, buds that become limbs, and pharyngeal pouches The tail remains in most adult vertebrates (humans lose it during fetal development) Only fish and immature amphibians retain pharyngeal pouches (in humans it develops into structures in the throat)
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Analogous structures Body part that is similar in function as a body part of another organism but is structurally different
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Structural patterns are clues to the history of a species
Vestigial structures: remnants of an organ or structure that functioned in an earlier ancestor Reduced in size and function
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Evolutionary biology today
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Molecular and genetic evidence support fossil and anatomical evidence
DNA sequence analysis Pseudogenes Homeobox genes Protein comparisons
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Biological Molecules As physical changes occur in the fossil record, genes that determine the species characteristics should have changed as well Proteins and DNA sequences can be used to determine the relatedness of species
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Evolution unites all fields of biology
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