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Sensation &Perception
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Sensation the immediate response in the brain caused by the stimulation of a sensory receptor Schacter’s definition: sensation = simple awareness due to the stimulation of a sense organ Perception the process of organizing sensory information into meaningful patterns
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Historical Background
Psychophisicists sought answers to sensory questions what is the weakest light that can be seen? what are the limits of the sense organs? how much change must occur in a stimulus before a sense organ will respond? attempted to quantitate relationship between actual physical properties of a stimulus and perceived properties of a stimulus
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Important Figures: Johannes Muller Ernst Heinrich Weber
Gustav Theodore Fechner Johannes Muller (1801 – 1858) Ernst HeinrichWeber (1795 – 1878) Gustav Fechner ( )
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Johannes Muller systematically remove organs and tested their response to various chemicals and stimuli Noted that all nerves carry the same basic message; electrical activity Derived the Law of Specific Nerve Energies All nerve fibers carry the same type of message Sensory information is specified by the particular nerve fibers that are active
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Natural Law of Psychology
Ernst Weber JND smallest magnitude of change in a stimulus that can be detected by a human Natural Law of Psychology "the amount of change needed to produce a JND is a constant proportion of the original stimulus" Weber's Fractions e.g. loudness fraction is 1/10
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If a radio is playing at volume "5", how loud must it get before you can tell that the volume has increased?
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Gustav Fechner "What is the range over which we can detect a JND?
led to determination of absolute threshold minimum amount of a stimulus that can be detected
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Subliminal Perception and Perceptual Defense
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Subliminal Perception
perception of a stimulus that is below the threshold for conscious recognition became part of "pop" psychology "utilized" by marketing and advertising made famous with the presentation of the film PICNIC in 1957
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Is subliminal perception a real phenomenon?
It can be "reproduced" to a limited degree in a laboratory under extremely precise conditions appears to have absolutely no effect on behavior particularly useless in attempting to influence choice and motivation
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Perceptual Defense resistance to perceiving threatening or disturbing stimuli became a part of "pop" psychology - also studied by legitimate scientists extensive investigations have dismissed the existence of perceptual defense
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Sensation and Perception
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General Principles of Sensory Coding
Stimulus Reception Transduction Coding
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Stimulus Reception Sensory Receptor
any external, physical energy detectable by a sensory receptor Reception the absorption of physical energy by a sensory receptor Sensory Receptor a specialized neuron that detects a particular category of physical energy
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Transduction conversion of the physical energy to an electrochemical pattern in the neurons Coding the one to one correspondence between the stimulus and the receptor activity
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The Visual System
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Stimulus the visible spectrum
the human visual system responds only to electromagnetic radiation of wavelengths between of nm
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Anatomy of the Visual System
eyes are suspended in two bony pockets called the orbits held in place and moved by three pairs of muscles
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The Parts Cornea Sclera translucent membrane that bends light inward
focuses light Sclera white opaque component provides protection and serves as an attachment for the external muscles
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Iris Crystalline Lens pigmented aperture of the eye
opens and closes to adjust the amount of light entering the eye Crystalline Lens located just behind the iris ciliary muscles adjust the shape of the lens so as to bend light and focus it on the fovea process called accommodation
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Retina Vitreous Humor thin layer lining the inside of the eye
receives light rays Vitreous Humor viscous jelly-like fluid that fills the inner compartment of the eye maintains shape and ocular pressure
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Common Visual Defects Myopia (nearsightedness)
eyeball is too long - image falls short of fovea requires concave lens to adjust Hyperopia (farsightedness) eyeball is too short - image falls beyond fovea requires convex lens to adjust Astigmatic Eye or Astigmatism misshapen cornea - part of image is in focus requires asymetrical lens to correct focus
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Presbyopia or Old Vision
farsightedness due to agin lens becomes less flexible with age treated with magnification requires bifocal lens of hyperopia or myopia also present
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Trace a photon of light through the eye.
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The Retina
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fovea centralis or just fovea
depressed spot in the macula which contains only cones rods are "shoved" aside point of highest visual acuity Why can't you see a dim star if you look directly at it?
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Optic Disk Locate your blind spot "blind spot"
ganglion cell axons exit the sclera to form the optic nerve no rods or cones (blind) Locate your blind spot
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The Photoreceptors Two general types Rods Cones
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Rods Cones named for long slender shape absent in fovea
numbers increase toward the periphery of the retina respond well to dim light bleached by bright light poor detail Cones named for pointed shape high density in the fovea mixed in periphery with rods respond poorly in dim light respond best in bright light detail vision is good
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Transduction photopigments consist of two parts opsin (protein)
retinal (a lipid)
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several different forms of the opsin protein e.g. Rhodopsin
pigment found in rods normally exists in the all-cis state
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when exposed to light the photogiment splits into its two component opsin and retinal
results in a change in the membrane which allows a signal to be sent to the brain
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The Visual Information Pathway
retinal ganglion cell axons form the optic nerve synapse in the Lateral Geniculate Nucleus (LGN) of the thalamus neurons from the thalamus send axons to primary visual cortex via the optic radiations some fibers synapse in the superior colliculus
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Where does information go after reaching occipital cortex?
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Coding How do we see color? Three Theories
The Trichromatic (Young Helmholtz) Theory The Opponent Process Theory The Retinex Theory
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The Trichromatic Theory
Color is coded or perceived as the result of variations in the "firing" rate of three types of cones short wavelength medium wavelength long wavelength
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evidence for the trichromatic theory comes from
Electrode recordings in primates color vision abnormalities in humans genetic defects in color vision are the result of anomalies in one or more of the three types of cones
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Protanopia and Deuteranopia
confuse red and green see the world in shades of yellow and blue red and green both look greenish to yellowish
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Normal Protanope Deuteranope
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Tritanopia third color defect extremely rare
equally represented in male and female populations cannot see short wavelengths lack blue receptor completely Tritanope Normal Deuteranope
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The Opponent-Process Thoery
Color is perceived in terms of paired opposites red vs green blue vs yellow goes beyond trichromatic theory - can explain the negative color afterimage phenomenon
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negative after image the image seen after a portion of the retina is exposed to an intense visual stimulus consists of colors which are complementary to those of the physical stimulus complementary colors are those that make white or gray when mixed together
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firing rate of ganglion cells appears to be most important mechanism producing negative afterimage
after being excited or inhibited for a long time, ganglion cells undergo a rebound effect fire faster or slower than normal
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Retinex Theory Retina and Cortex = Retinex
developed to explain the phenomenon of color constancy despite the fact that the wavelength of stimulus is changing, we perceive the colors as constant
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Artificial light gray filter outdoors daylight
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How do we impose meaning on what we see?
One way is through the process of perceptual constancy The knowledge that the physical characteristics of objects remain constant even though the image reaching the retina has changed.
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Size and Shape constancy
perceived size and shape of physical objects remains constant even though retinal image is different or changing requires experience - learned e.g. experience of Dr. Colin Turnbull’s and Pygmy tribe
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Brightness constancy refers to the fact that the apparent brightness of an object remains constant under changing light conditions but our experience can "fool" us into seeing lighting differences where there are none
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clues we use to create meaningful patterns
Perceptual Grouping clues we use to create meaningful patterns
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Proximity Similarity Continuity
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Closure Common region
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common fate or movement
Contiguity (occurring close in time) - basis for drawing causal conclusions can lead to erroneous causal conclusions e.g. thunder and lightening
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(both innate and learned cues)
Depth Perception (both innate and learned cues) Innate Accommodation (monocular clue) the lens of the eye adjusts its shape to focus on objects receptors in ciliary muscles provide information about the distance of an object works within about 4 feet of the eyes Convergence (binocular) eyes turn inward to focus on near objects ciliary muscles send info to brain Retinal Disparity (binocular) approx. 6 cm distance between eyes (retinas) slightly different images on retina fusing images creates 3D vision
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Learned Pictorial Cues
Linear Perspective convergent parallel lines Relative Size requires knowledge of size constancy Height in the Picture Frame drawing an item closer to the horizon line makes it appear more distant Light and Shadow in the "real" world thre are visual differences between light and shadow creating light and shadow in 2D space gives a 3D perception
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Overlap or Interposition Texture Gradient
a distant object which partially blocks another object is perceived as closer Texture Gradient distant textures look smoother than near textures Aerial Perspective adding haziness or blurriness to objects make them appear more distant Motion Parallax imagine riding down a highway in a car - near objects appear to pass in front of more distant objects
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The Auditory System
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Sound Waves Rhythmic movement of air molecules A series of compressions (peaks) rarefractions (valleys) Characteristics of sound waves Pitch (high vs low tone) Corresponds to frequency or waves per second Loundness Corresponds to the amplitude of the wave
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Transduction Pinna Outer ear
Funnels sound waves through external canal to tympanic membrane Tympanic Membrane Sound waves press against it moving it in and out Auditory Ossciles (middle ear) Movement of tympanic membrane sets up vibrations in malleus Malleus taps against incus Incus attached to stapes Stapes contacts oval window at the opening to the scala tympani Oval Window Begins to move in and out setting up movement of the perilymphatic fluid in the cochlea
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Cochlea Snail shaped organ of hearing Contains three fluid filled chambers Scala vestibuli Scala media Scala tympani Organ of Corti Located in the scala media Consists of Basilar membrane Tectorial membrane Hair cells Movement of endolymphatic fluid causes displacement of basilar membrane Hair cells embedded in basilar membrane have stereocilia that stick up through the tectorial membrane Displacement of basilar membrane results in shearing forces of the stereocilia which causes Na+ channels to open
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Axons of the hair cells form the auditory division of the 8th
Axons of the hair cells form the auditory division of the 8th. cranial nerve Information is sent to Cochlear nuclei in the brain stem Inferiror colliculus Thalamus Primary Auditory Cortex in the temporal lobe
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Anatomical or Place Coding
Coding Pitch Anatomical or Place Coding Different areas of the basilar membrane are stimulated by different sound frequencies True for medium to high pitch (300 – 20,00 Hz)
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Temporal or Frequency theory
For sound of < 200 Hz or less there is no corresponding place on the basilar membrane Sounds < 200 Hz produce a synchronous firing pattern in the basilar membrane Volley Theory A modification of temporal or frequency theory Nerve fibers work as a “squad” Group may not respond to every wave but to every 3rd. or 4th. Etc. as a group (a volley)
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