1. Bankfull discharge (Q bkf ), an important concept –Fills the channel and does work on boundaries –"Dominant discharge” Greatest total volume of sediment moved Frequency-Magnitude Concept (Fig. 3.10, text) For suspended: Q d is ~Q bkf a “channel forming flow” –Peak discharge that occurs on average once in 1.5 - 2 years –Range 1-10 years, depending on channel type –Applies best to gravel-cobble rivers with floodplains Magnitude sediment load Freq. of Q SuspendedBedload The highest flows are not the “dominant discharge”!

2. MORPHOLOGIC CLASSIFICATIONS OF STREAMS Based solely upon physical features and processes 1) Reach scale (Rosgen 1994) based on physical appearance (hydrogeomorphic processes implicit) 41 channel types based on sinuosity, width to depth ratio, confinement, slope repeatable and widely- applicable (but "cook- book”) used by agencies for channel (physical) restoration Easily misapplied (and often fails?)

3. 2) Reach scale (Montgomery and Buffington 1997) -focus on processes that form and maintain channels Increasing stream gradient 5 types of channels (for mountain streams) -cascade -step-pool -plane-bed -pool-riffle -sand-bed

4. 3) Hierarchical, multi-scale -Processes acting at LARGE scales influence/constrain patterns at small scales -For example: -basin geology constrains the kind of channel reach morphology and habitat sediment types -Climate determines flood frequency and stability of riffle gravels over an annual cycle -Hierarchical classification systems allow choice of the level of habitat resolution that is required to meet specific objectives

5. Frissel et al. 1986 Watershed  Segment  Reach  Pool/riffle  Microhabitat Spatial and Temporal Scale Relationships

6. Other Hierarchical Classifications –Hawkins et al. (1993) Classify “channel geomorphic units”

7. Substrate-Organism Relationships Substrate = Substratum (Substrata) Fundamental Roles –1) Habitat –2) Food –3) Protection

8. Ways to Characterize Substrate 1) type 2) size (area) and shape 3) heterogeneity 4) texture 5) stability

9. Substrate Type & Size Inorganic -- mineral –Exposed Bedrock –Particles (from weathered bedrock) Wentworth Scale: B > C > G > Sa > Si > C Organic –Fine Particulate OM (FPOM, < 1mm) –Course Particulate OM (e.g., whole leaves) (CPOM, > 1mm) –Wood –Living plant surfaces / Moss

10. Mineral particles –SIZE [Table 12.3 b (Minshall)] Richness (# spp.) –increases Species density (#spp./m 2 ) –declines Abundance (#individuals) –increases Density (# ind./m 2 ) –declines … a bit –Same pattern in Table 12.3a –SHAPE [Table 12.3a] Irregular shape more microhabitats

11. Mineral Particle Mixtures Table 12.1 (Minshall) –Density and biomass of invertebrates Sand supports low abundance and biomass Rubble (pebble + cobble) and gravel support more Sand Gravel-Cobble Cobble-Boulder Interstitial spaces between substrate particles influence many aspects of habitat quality –Habitat space –Flow (Fig.8-12, Ward) –Oxygen –Stability

12. Fish spawning –Salmonids can create redds in gravel having median size up to 10% of their body length –Particles < 1 mm reduce gravel permeability and impair egg development (reduced flow and oxygen) –Particles 1-10 mm block intergravel pores and impede “swim-up” of newly-hatched fish.

13. Mineral vs. Organic substrates –Table 3.4 -- classic study sand supports fewest numbers and species organic substrates support more (but they are seasonal) Wood –Steep channels: creates habitat (pools) and increases CPOM retention. –Low gradient channels with silt/sand bottoms: stable habitat Table 3.5 Satilla River, GA

14. Surface Texture Scale relative to the organism Field observation - more “pits”, more algae or invertebrates Experimental approaches: –Example 1: Algae (Fig. 2, Bergey) 4 surfaces with different “pittedness” Experimental sand abrasion –Interpret graph Effect of abrasion level on diatom density? Effect of surface ”pittedness” on density? (panel A vs. B)

15. Stability –Particle resistance to movement by high flows Critical erosion velocity (Fig. 1.7) –Hydro-Geomorphic setting Slope, channel confinement difference in Q regime among streams –Fig. 4.10 – Moss on stones ? ? A rolling stone gathers no moss … the proof!

16. Heterogeneity Generally, RIFFLES > POOLS for both benthic Diversity and Production. WHY? More substrate diversity Cob BoulPbl Gvl

17. Heterogeneity a) Particle sorting by flowing not perfect. Many “patches” (range) of particle sizes. b) Scales of heterogeneity –Local patch : some terms Embeddeness - excess fines (draw picture) –Channel Armoring - loss of fines(draw picture) mid-channel (thalweg) to edges –Reach scale riffles vs. pools –Whole river system uplands channels more heterogeneous than lowland (excluding floodplains)

18. Organism-substrate relationships? Many loose patterns -- interacting factors, due to correlations among important variables: Current Oxygen Food Stability –Riffles: coarse substrate mixtures with high interstitial flows, high O 2, trapped CPOM, and relatively high stability –Pools: fine substrates, low interstitial flows, low O 2 ; lots of organic matter, low stability