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Leaves Chapter 7 Copyright © McGraw-Hill Companies Permission

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1 Leaves Chapter 7 Copyright © McGraw-Hill Companies Permission
Required for Reproduction or Display

2 Outline Overview Leaf Arrangements and Types Internal Structures of Leaves Stomata Mesophyll and Veins Specialized Leaves Autumnal Changes in Color Abscission Relevance of Leaves

3 Overview All leaves originate as primordia in the buds.
At maturity, most leaves have a stalk (petiole) and a flattened blade (lamina) with a network of veins (vascular bundles). Leaves of flowering plants are associated with leaf gaps and have an axillary bud at the base, with or without a stipule. May be simple (single blade) or compound (divided into leaflets).

4 Leaf Anatomy Simple Leaf “Erguvan” Dişbudak Compound Leaf

5 elliptical ovate lanceolate cordate spatulate pinnately lobed Dişbudak
Kızılcık cordate spatulate pinnately lobed Erguvan Meşe

6 Gymnosperm Leaves awl-like scale-like ARDIÇ

7 Overview Pinnately compound leaves have leaflets in pairs along the rachis, while palmately compound leaves have all the leaflets attached at the same point at the end of the petiole. Pinnately compound leaves may be further subdivided an thus be referred to as bipinnately compound.

8 Leaf Arrangements and Types
Leaves are attached to stems at nodes, with stem regions between nodes known as internodes. Phylotaxy (leaf arrangement) generally occurs in one of three ways: Alternate Opposite Whorled

9 Bipinnately compound twice pinnate, the primary leaflets are divided into secondary leaflets, also written 2-pinnate Gleditsia triacanthos Gladiçya

10 Leaf Arrangements and Types
Arrangement of veins in a leaf or leaflet blade may also be pinnate or palmate. Pinnately veined leaves have a main midvein within a midrib. Secondary veins branch from midvein. Palmately veined leaves have several primary veins that fan out from the base of the blade. Parallel in monocots Dichotomous in Ginkgo Karaağaç Sığla

11 Overview Green leaves capture sunlight and thus go through photosynthesis. Lower surfaces of leaves are dotted with stomata which allow carbon dioxide to enter and oxygen and water to diffuse out. Guard Cells control stomatal opening. Transpiration occurs when water evaporates from the leaf surface. Guttation - Root pressure forces water out hydathodes.

12 Internal Structure of Leaves
Epidermis is a single layer of cells covering the entire surface of the leaf. Upper epidermal cells are devoid of chloroplasts. Waxy cuticle often present. Different glands may also be present in the epidermis.

13 Stomata Lower epidermis of most plans is perforated by numerous stomata. Guard cells originate from the same parent cell, and contain chloroplasts. Primary function includes regulating gas exchange between leaf interior and the atmosphere, and the evaporation of water. Cell water pressure regulates guard cells which in turn regulate stomata.

14 Mesophyll and Veins Most photosynthesis takes place in the mesophyll (chlorenchyma) between the two epidermal layers. Palisade Mesophyll - Uppermost layer Contain most of leaf’s chloroplasts. Spongy Mesophyll - Lower layer Veins (Vascular bundles) are scattered throughout the mesophyll. Consist of xylem and phloem tissues surrounded by the bundle sheath.

15 Monocot leaves Bulliform cells Usually no mesophyll differenetiation
Midrib Usually no mesophyll differenetiation Bulliform cells (either side of the midrib) have thin walls and collapse under dry conditions causing folding or Rolling.

16 Specialized Leaves Shade Leaves
Leaves in the shade receive less total light, thus tend to be thinner and have fewer hairs than leaves on the same tree exposed to direct light. Leaves of Arid Regions Many have thick, leathery leaves and few stomata or sunken stomata. Some have succulent, water-retaining leaves, or dense, hairy coverings. Pine trees have sunken stomata, thick cuticle and a layer of thick-walled cells (hypodermis) beneath the epidermis.

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18 Specialized Leaves Tendrils
Modified leaves that curl around more rigid objects helping the plant to climb or support weak stems. Become coiled like a spring as they develop. When contact is made, the tip curls around the object, and the direction of the coil reverses.

19 S T P Specialized Leaves Spines, Thorns, and Prickles
Spines - Modified leaves designed to reduce water loss and protect from herbivory. Thorns - Modified stems arising in the axils of leaves of woody plants. Prickles - Outgrowths from the epidermis or cortex. S T P

20 Specialized Leaves Storage Leaves – Succulent desert plants, onion, lilly and other bulbs Flower-Pot Leaves - Urn-Like Pouches Flower-pot leaf -Dischidia

21 Window Leaves - Leaves buried in ground.
Specialized Leaves Window plant-Fenestraria Window Leaves - Leaves buried in ground. Reproductive Leaves - New plants at tips. Floral Leaves - Bracts Air plant-Kalanchoe

22 Insect-Trapping Leaves Pitcher Plants Sundews
Specialized Leaves Insect-Trapping Leaves Pitcher Plants Sundews Copyright © McGraw-Hill Companies Permission Required for Reproduction or Display Copyright © McGraw-Hill Companies Permission Required for Reproduction or Display

23 Insect-Trapping Leaves Venus’s Flytraps Bladderworts
Specialized Leaves Insect-Trapping Leaves Venus’s Flytraps Bladderworts Copyright © McGraw-Hill Companies Permission Required for Reproduction or Display Copyright © McGraw-Hill Companies Permission Required for Reproduction or Display

24 Leaf colors

25 Autumnal Changes in Leaf Color
Cholorplasts of mature leaves contain several groups of pigments. Chlorophylls - Green Carotenoids - Yellows In fall, chlorophylls break down and other colors are revealed. Water soluble anthocyanins (red:Acidic or blue:Alkaline, intermediate shades: neutral) and betacyanins (red) may also be present in the vacuole.

26 Abscission Deciduous plants drop their leaves seasonally. Occurs as a result of changes in an abscission zone near the base of the petiole of each leaf. Cells of the protective layer become coated and impregnated with suberin. Cells of the separation layer divide and become gelatinous. Pectins in the middle lamella broken down. Separation becomes easy.

27 Leaf Abscission Zone

28 Human and Ecological Relevance of Leaves
Landscaping: cooling+aesthetics Food: cabbage, parsley, lettuce, oregano peppermint etc. Dyes: yellow;Bearberry, reddish;Henna Ropes and Twine: Agave, Sansevieria, Panama Hat Palm Fuel: Yareta leaves contain resins Drugs Tobacco Marijuana Cocaine Aloe vera Brewery:Tequila from agave, various teas from relatives of garden camellia Insecticides: Mexico’s cockroach plant, India’s Neem tree are powerfull insecticides Waxes: Carnauba and caussu wax from tropical palms

29 Copyright © McGraw-Hill Companies Permission Required for Reproduction or Display

30 Human and Ecological Relevance of Leaves

31 Human and Ecological Relevance of Leaves
Agave

32 Human and Ecological Relevance of Leaves
Atropa belladonna

33 Human and Ecological Relevance of Leaves
Bearberry-yellow dye

34 Human and Ecological Relevance of Leaves
Digitalis purporea Coca

35 Human and Ecological Relevance of Leaves
Marijuana- Tetrahydrocannabinol (THC)

36 Human and Ecological Relevance of Leaves

37 Human and Ecological Relevance of Leaves

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39 Review Overview Leaf Arrangements and Types Internal Structures of Leaves Stomata Mesophyll and Veins Specialized Leaves Autumnal Changes in Color Abscission Relevance of Leaves


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