Fig. 1 Mean swarm response to an oscillating swarm marker.

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Fig. 1 Mean swarm response to an oscillating swarm marker. Mean swarm response to an oscillating swarm marker. (A) Trajectories (>40 s long) of individual midges (each color corresponding to a different midge) are individually convoluted but remain localized over the ground-based swarm marker (black square). (B) Sketch of our experimental setup. Swarms form inside a plexiglass cube measuring 122 cm on a side and are imaged using three cameras mounted outside the enclosure. The swarm marker (in dark gray) is mounted on a linear stage (in red) that can be oscillated over a range of controlled frequencies and amplitudes along the direction indicated by the white arrows, which we label as the x direction. z increases vertically from the swarm marker (antiparallel to gravity), with the marker itself at z = 0. Midge development tanks (light blue) and four infrared light-emitting diode arrays (yellow; additional arrays on top of the enclosure are not shown) are also shown. (C) Phase-averaged position of the center of the swarm marker XM and the center of mass of the swarm XS. The swarm center of mass tracks the sinusoidal motion of the marker, although with a reduced amplitude and a phase lag. (D) The amplitude of the swarm center-of-mass motion AS as a function of the amplitude of the marker motion AM for two different oscillation frequencies, showing a linear relationship between the two. The shaded area shows the SEM. Kasper van der Vaart et al. Sci Adv 2019;5:eaaw9305 Copyright © 2019 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works. Distributed under a Creative Commons Attribution NonCommercial License 4.0 (CC BY-NC).