~ 5,000 aquatic beetles worldwide Coleoptera ~ 5,000 aquatic beetles worldwide
Most live in substrate: Adults usually leave water temporarily - dispersal. One time - Elmidae (riffle beetles) Several times - Dytiscidae, Hydrophilidae
Respiration? 1. Self-contained air reserves (Dytiscidae - diving beetle) 2. Transcuticular respiration (gills in larvae) 3. Plastron respiration (adult Dryopidae) 4. Piercing plant tissues (larval Chrysomelidae) Slender antennae Hind coxae extend posteriorly divide first abdominal segment Swimming hairs on legs
Respiration? 1. Self-contained air reserves (Dytiscids) 2. Transcuticular respiration (gills in larvae) 3. Plastron respiration (adult Dryopidae) 4. Piercing plant tissues (larval Chrysomelidae)
Respiration? 1. Self-contained air reserves (Dytiscids) 2. Transcuticular respiration (gills in larvae) 3. Plastron respiration (adult Dryopidae) 4. Piercing plant tissues (larval Chrysomelidae) Long toed water beetles Woolly hairs except last abdominal segment Short antennae
Respiration? 1. Self-contained air reserves (Dytiscids) 2. Transcuticular respiration (gills in larvae) 3. Plastron respiration (adult Dryopidae) 4. Piercing plant tissues (larval Chrysomelidae)
Varied life histories Some larvae occur year round, adults for short period (Psephenidae) Water pennies Larvae are aquatic, adults terrestrial
Varied life histories Others occur as larvae and adults year round – Elmidae = riffle beetles Long antennae 5-segmented tarsi Crawl on bottom - do not swim = no swimming hairs on hind legs 5-6 abdominal segments
Varied life histories Others have long adult; larva only around a few weeks - Hydrophilidae Water scavenger beetles Adult 3 segmented club antennae 5 segmented tarsi Larvae Large mandibles 4 segmented legs
Scirtidae = marsh beetles Adults = terrestrial! Scirtidae = marsh beetles Small (5 - 15 mm) Larvae have long antennae 4-segmented legs, single claw
Beetle pupae are mostly terrestrial Under stones or logs Few marine beetles - substrate dwellers
Larval morphology = diverse Sclerotized head capsule Mandibles, maxillae, labium Gill-like appendages in some (Gyrinidae = whirligig beetles)
Coleoptera
Coleoptera
Coleoptera
Coleoptera
The key 2 compound eyes that appear to be divided = Gyrinidae, whirligig beetles
Eyes not divided, hind coxae divided into plates that cover abdominal segments 1-2, or 3 Haliplidae, crawling water beetles
Hind coxae not expanded into plates, divide abdominal segment 1 Dytiscidae, predaceous diving beetles
Hind coxae not dividing abdominal segment 1, antennae clubbed with cuplike segment at base Hydrophilidae, water scavenger beetles
Antennae short with comb-like club, body length 5-6.5 mm Dryopidae, long-toed water beetles
Antennae w/o comblike club, body length < 4.5 mm Elmidae, riffle beetles