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Chapter 22 Descent with modification: A Darwinian view of life.

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1 Chapter 22 Descent with modification: A Darwinian view of life

2 Natural Selection: A Summary Individuals with certain heritable characteristics survive and reproduce at a higher rate than other individuals Natural selection increases the adaptation of organisms to their environment over time If an environment changes over time, natural selection may result in adaptation to these new conditions and may give rise to new species Video: Seahorse Camouflage Video: Seahorse Camouflage Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings

3 Fig. 22-12 (b) A stick mantid in Africa (a) A flower mantid in Malaysia

4 Anatomical and Molecular Homologies Homologous structures are anatomical resemblances that represent variations on a structural theme present in a common ancestor Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings

5 Fig. 22-17 Humerus Radius Ulna Carpals Metacarpals Phalanges HumanWhale Cat Bat

6 Convergent Evolution Convergent evolution is the evolution of similar, or analogous, features in distantly related groups Analogous traits arise when groups independently adapt to similar environments in similar ways Convergent evolution does not provide information about ancestry Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings

7 Fig. 22-20 Sugar glider Flying squirrel AUSTRALIA NORTH AMERICA

8 Variation Between Populations Most species exhibit geographic variation, differences between gene pools of separate populations or population subgroups

9 Fig. 23-3 13.1719XX10.169.12 8.11 1 2.4 3.145.18 67.15 9.10 12.19 11.1213.17 15.18 3.84.165.14 6.7 XX

10 Chapter 23 The evolution of populations

11 Hardy-Weinberg equilibrium describes the constant frequency of alleles in such a gene pool If p and q represent the relative frequencies of the only two possible alleles in a population at a particular locus, then – p 2 + 2pq + q 2 = 1 – where p 2 and q 2 represent the frequencies of the homozygous genotypes and 2pq represents the frequency of the heterozygous genotype

12 Fig. 23-7-1 Sperm C R (80%) C W (20%) 80% C R ( p = 0.8) C W (20%) 20% C W ( q = 0.2) 16% ( pq ) C R C W 4% ( q 2 ) C W C W C R (80%) 64% ( p 2 ) C R C R 16% ( qp ) C R C W Eggs

13 Genetic Drift The smaller a sample, the greater the chance of deviation from a predicted result Genetic drift describes how allele frequencies fluctuate unpredictably from one generation to the next Genetic drift tends to reduce genetic variation through losses of alleles Animation: Causes of Evolutionary Change Animation: Causes of Evolutionary Change

14 Fig. 23-9 Original population Bottlenecking event Surviving population

15 Gene Flow Gene flow consists of the movement of alleles among populations Alleles can be transferred through the movement of fertile individuals or gametes (for example, pollen) Gene flow tends to reduce differences between populations over time Gene flow is more likely than mutation to alter allele frequencies directly

16 Gene flow can decrease the fitness of a population In bent grass, alleles for copper tolerance are beneficial in populations near copper mines, but harmful to populations in other soils Windblown pollen moves these alleles between populations The movement of unfavorable alleles into a population results in a decrease in fit between organism and environment

17 Directional, Disruptive, and Stabilizing Selection Three modes of selection: – Directional selection favors individuals at one end of the phenotypic range – Disruptive selection favors individuals at both extremes of the phenotypic range – Stabilizing selection favors intermediate variants and acts against extreme phenotypes

18 Fig. 23-13 Original population (c) Stabilizing selection (b) Disruptive selection (a) Directional selection Phenotypes (fur color) Frequency of individuals Original population Evolved population

19 Heterozygote advantage occurs when heterozygotes have a higher fitness than do both homozygotes Natural selection will tend to maintain two or more alleles at that locus The sickle-cell allele causes mutations in hemoglobin but also confers malaria resistance Heterozygote Advantage

20 Fig. 23-17 0–2.5% Distribution of malaria caused by Plasmodium falciparum (a parasitic unicellular eukaryote) Frequencies of the sickle-cell allele 2.5–5.0% 7.5–10.0% 5.0–7.5% >12.5% 10.0–12.5%

21 Chapter 24 The origin of species

22 Conditions on early Earth made the origin of life possible Chemical and physical processes on early Earth may have produced very simple cells through a sequence of stages: 1. Abiotic synthesis of small organic molecules 2. Joining of these small molecules into macromolecules 3. Packaging of molecules into “protobionts” 4. Origin of self-replicating molecules

23 Protobionts Replication and metabolism are key properties of life Protobionts are aggregates of abiotically produced molecules surrounded by a membrane or membrane-like structure Protobionts exhibit simple reproduction and metabolism and maintain an internal chemical environment

24 The First Eukaryotes The oldest fossils of eukaryotic cells date back 2.1 billion years The hypothesis of endosymbiosis proposes that mitochondria and plastids (chloroplasts and related organelles) were formerly small prokaryotes living within larger host cells An endosymbiont is a cell that lives within a host cell

25 Fig. 25-9-4 Ancestral photosynthetic eukaryote Photosynthetic prokaryote Mitochondrion Plastid Nucleus Cytoplasm DNA Plasma membrane Endoplasmic reticulum Nuclear envelope Ancestral prokaryote Aerobic heterotrophic prokaryote Mitochondrion Ancestral heterotrophic eukaryote

26 Aneuploidy Aneuploidy is an abnormal number of chromosomes, and maybe detrimental Polyploidy are organisms containing more than two paired (homologous) sets of chromosomes.

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28 Genetic Variation Variation in individual genotype leads to variation in individual phenotype Not all phenotypic variation is heritable Natural selection can only act on variation with a genetic component

29 Fig. 23-2 (a) (b)

30 Unit 4 C22 45&46, 70&71, 79&80C23 5&6, 11&12, 31&32, 44&46, 50&52, 58&60, 69&70, 89&90C24 14&18C25 5&12, 54&60, 93&screen shot


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