1. Senses: Taste and Smell Chemical “conversation” – Especially important for large social groups – Recognize territory (Dog) – Navigate during migration (Salmon)

2. Continued Gustation- taste Olfaction- smell Both are dependent on chemoreceptors Land animals – taste is detection of chemicals within a solution – Smell is a detection of chemicals that travel through air No difference in aquatic animals

3. Taste in Humans Taste Buds- “modified epithelial cells organized” Papillae 4 basic tastes- sweet, sour, salty, bitter – Extra one umami (Japanese for delicious) Elicited by Glutamate – Complex flavors = Brain integrating various inputs from receptors

4. Continued Chemoreceptors have channels present within their plasma membranes that ions diffuse through. Salt = Na + Sourness = H + Bitterness = K + Ions diffuse into the cell – cell depolarizes Depolarization causes cell to release neurotransmitter on a sensory neuron

5. Smell in Humans Olfactory receptor cells- neurons in nose – Send impulses to the olfactory bulb of the brain Odorant receptors (specific proteins) that get bind to the odorous substances that diffuse into the nose – Signal transduction pathway opened G protein, enzyme adenylyl cyclase, 2 nd messenger cyclic AMP – 2 nd messenger cyclic AMP opens channels in the plasma membrane that are permeable to both Na + and CA 2+ ions – Influx in ions depolarizes the membrane = ACTIVE POTENTIAL

6. Continued More than a 1000 Ors= 3% of human genes Receptors in Brain for Taste and Smell are independent yet they interact.

7. Vision in Invertebrates Ocellus- provides information about information about light intensity and direction – Doesn’t create images Compound eyes (insects) – Contain several thousand light detectors= ommatidia Each has its own focusing lens Humans can see flashes that occur at 50 flashes per second, while insects can see them at 330 times per second

8. Vision in Human Photoreceptors – in retina Absorb light send info to brain through optic nerve Two types – Rod cells – Cone cells

9. Differences Rod cells – More sensitive to light, function better in dim light – absorb all wavelengths= monochrome vision – cone cells provide better visual acuity – widely dispersed throughout the cell= wider vision Cone cells – Less sensitive to light, function better in bright light – sensitive to red, green, and blue= color vision – concentrated near the fovea= acute area of vision

10. Structure of Eyes Sclera- White outer layer Choroid- pigmented inner layer Conjunctiva- mucous membrane Cornea- lets light in and acts as a fixed lens Iris- regulates amount of light Pupil- hole in center of the iris Retina- inner most layer contains photoreceptors

11. Processing Visual Information Convergence – Bipolar cells (synapses between either rods and cones and neurons) Combine with impulses and pass them to ganglion cells (sensory neurons on the optic nerve) Edge enhancement – Ganglion cell is stimulated by light on the circle on the retina called the receptive field. 2 types Light falls on the center of the receptive field but also the periphery= stimulation reduced. Light falls on periphery and falls on the center of the receptive field= stimulation is reduced Herman Gird- looks whiter next to black area

12. Continued Contralateral Processing – Left and right optic nerves meet at optic chiasma. – Impulses cross over to opposite optic nerves – Allows for brain to deduce distance and size