1. The northern goshawk as an example Desired Conditions: Habitat, Biodiversity, and Foodwebs

2. Morphological and Behavioral Characteristics Adapted to forest environments Adapted to forest environments Long tail, short wings Long tail, short wings Sub-canopy forager Sub-canopy forager Primarily perch hunter Primarily perch hunter Short-sit, short-flight hunting behavior Short-sit, short-flight hunting behavior Searches for prey in low vegetation column Searches for prey in low vegetation column Broad diet Broad diet

3. Foraging Habitat Occupies many forest types Occupies many forest types Forest structure Forest structure –Lifted crowns (mature forests) –Open understory (flight space) –Hunting perches Strong evidence of food limitation Strong evidence of food limitation

4. Birds American robin American robin Band-tailed pigeon Band-tailed pigeon Blue grouse Blue grouse Mourning dove Mourning dove Steller’s jay Steller’s jay Northern flicker Northern flicker Hairy woodpecker Hairy woodpecker Mammals Chipmunks Chipmunks Cottontail Cottontail Jackrabbit Jackrabbit Mantled ground squirrel Mantled ground squirrel Red squirrel Red squirrel Abert’s squirrel Abert’s squirrel Prey

5. Kaibab Goshawk Diet % frequency% biomass

6. Proportion of Pairs Breeding Pairs Breeding (%) Year

7. Prey Density and Goshawk Productivity Prey Density ha -1 Year Goshawk productivity per Territory

8. Precipitation Goshawk breeding Annual precipitation and goshawk breeding 101 years precipitation Kaibab Plateau

9. Habitat Synthesis: The Goshawk Food Web

10. Management Recommendations for the Northern Goshawk in the Southwestern United States Richard T. Reynolds Richard T. Reynolds Russel T. Graham Russel T. Graham M. Hildegard Reiser M. Hildegard Reiser Richard L. Bassett Richard L. Bassett Patricia L. Kennedy Patricia L. Kennedy Douglas A. Boyce, Jr. Douglas A. Boyce, Jr. Greg Goodwin Greg Goodwin Randall Smith and Randall Smith and E. Leon Fisher E. Leon Fisher

11. GOSHAWK HABITATGOSHAWK PREY SPECIES Composition & structure Tree longevity Sustainability Disturbance factors Landscape patterns/scale Goshawk nesting habitat Goshawk foraging habitat Species 1 Foods & Habitat Species 2 Foods & Habitat Species 3 Foods & Habitat Species 4 Foods & Habitat FHE SYNTHESIS OF FOREST HABITAT ELEMENTS (FHE) SYNTHESIS OF FOREST HABITAT ELEMENTS WITH FOREST ECOLOGY SUSTAINING LANDSCAPE MIX OF FOREST HABITATS FOREST ECOLOGY

12. Habitat, biodiversity, foodwebs Trees grouped with interlocking crowns Trees grouped with interlocking crowns Grass-forb-shrub openings between tree groups Grass-forb-shrub openings between tree groups All age classes and as much old forest as is ecologically sustainable All age classes and as much old forest as is ecologically sustainable High interspersion of age classes High interspersion of age classes

13. Habitat, biodiversity, foodwebs Trees grouped with interlocking crowns

14. Habitat, biodiversity, foodwebs As much old forest in groups as is ecologically sustainable

15. Habitat, biodiversity, foodwebs High age-class interspersion

16. Habitat, biodiversity, foodwebs Large tree component provides: –Snags –Logs –Woody debris

17. Habitat, biodiversity, foodwebs grass/forb/shrub

18. Poles 25-80 years Yellow pine 200+ years Blackjack 80-200 years Grass/ seedlings Scale - feet 100 200 300 0 ”Historical” conditions Fort Valley Experimental Forest Desired conditions provide habitats for goshawk food web

19. Unique elements Multi-species focus, not single species!! Multi-species focus, not single species!! Food web approach Food web approach –Information across trophic levels Desired conditions are: Desired conditions are: –Ecosystem specific, different DCs for different forest types

20. Unique Elements Landscape ecology Landscape ecology –Multiple goshawk home ranges = landscape scale –1,000-year planning horizon incorporates forest growth and succession –Spatial & temporal shifting mosaic of habitats Forest Restoration Forest Restoration –Natural composition, structure, and pattern of forest use as guides for assembling habitats –Restores natural disturbances, processes, and forest health (productivity)

21. Summation Small groups of trees with interlocking crowns Small groups of trees with interlocking crowns Scattered single trees Scattered single trees Grass-forb-shrub open interspaces between groups Grass-forb-shrub open interspaces between groups Snags, logs, woody debris Snags, logs, woody debris Spatial and temporal distribution of the above Spatial and temporal distribution of the above